Table 3. Diversity analyses of the Venezuelan and the Caribbean echinoid fossil record. ABBREVIATIONS: N – absolute number of genera; MDS – mean standing diversity without singletons; MDS+s – mean standing diversity with singletons; BC – boundary crossers.
 
 

 

Venezuela
Caribbean
Greater Antilles
Cuba
Jamaica
Lesser Antilles
Central America
 

 

N
MSD
MSD+
BC
N
MSD
MSD+s

N
MSD
MSD+s
B
N
MSD
MSD+s
B
N
MSD
MSD+s
B
N
MSD
MSD+s
B
N
MS
MSD+s
B
 

 





























  Early Cretaceous
9
2
3.67
0
41
21
23.67
34
25
15.5
15.5
25
20
12
12
20
5
4
4
5
1
0.5
0.5
1
19
7
8.67
14
  Late Cretaceous
0
0
0
0
51
24
29
18
45
18
23
16
39
17.5
21.5
15
9
3.5
4.5
2
1
0.5
0.5
0
14
7.5
7.5
1
  Paleocene
1
0
0.33
0
18
18
18
18
16
16
16
16
15
15
15
15
2
2
2
2
0
0
0
0
1
1
1
1
  Early Eocene
4
2
2
4
60
38.5
38.83
57
55
38
38.33
54
38
26
26.33
37
31
16.5
16.5
31
7
3.5
3.5
7
2
1.5
1.5
2
  Middle Eocene
5
4
4.33
4
65
61.5
61.5
63
60
20.5
20.83
60
45
41
41
44
34
31
31
30
10
8.5
8.5
8
2
1.5
1.5
1
  Late Eocene
13
4
6.33
4
76
50.5
52.83
36
67
45
45.67
29
55
35
37.67
27
33
20.5
20.83
9
18
7.5
9.17
7
1
1
1
1
  Early Oligocene
6
5
5
6
47
42
42
47
40
35
35
40
36
31.5
31.5
36
11
10.5
10.5
10
11
9
9
11
2
1.5
1.5
2
  Late Oligocene
6
1
1
6
54
40.5
42.83
32
48
31.5
34.17
21
45
28
30.67
19
11
10.5
10.5
9
11
11
11
11
2
2
2
2
  Early Miocene
15
9.5
10.17
14
42
34
34.67
37
26
23.5
23.5
24
21
18
18.33
18
16
12.5
12.5
14
18
10.5
12.5
10
14
8
8
13
  Middle Miocene
21
13
15
13
40
36
36.67
33
26
25
25.33
24
19
18
18.33
18
15
15
15
14
10
9.5
9.5
9
14
13
13.33
13
  Late Miocene
16
11
11
9
35
27
27.33
20
25
18.5
18.5
12
18
10.5
10.5
5
15
13.5
13.5
8
9
8.5
8.5
8
15
8
8.33
2
  Early Pliocene
11
9.5
9.83
10
22
20.5
20.83
21
12
12
12
12
5
5
5
5
9
9
9
8
8
8
8
8
3
2.5
2.5
3
  Late Pliocene
11
9.5
9.5
9
22
21.5
21.5
22
12
12
12
12
5
5
5
5
9
9
9
8
8
7.5
7.5
7
3
2.5
2.5
2
  Early Pleistocene
9
7
7
5
23
21
21
19
14
13
13
13
5
5
5
4
12
10.5
10.5
10
9
8
8
9
2
2
2
2
  Late Pleistocene
5
5
5

19
14.5
14.5

14
8.5
8.5

4
3.5
3.5

11
5.5
5.5

9
6.5
6.5

2
1.5
1.5

  Total Genera
51
157
122
102
54
36
38