TABLE 1. Meristic counts of all studied specimens of †Paralates . Numbers not in bold are subject to some uncertainty.

  Oligocene of Rouffach, no. Ruf Upper Eocene of Isle of Wight
(Chapelcorner Fish Bed)
Specimen # 6a/b 9_1 9_2 11/13_2 15a/b 18 P 59784 P 59785 P 59786 C 23632 C 23633
First dorsal fin (D1) - - >V - VII VII VII VII VII - -
Second dorsal fin (D2) I,10 I,9 I,9 I,9 I,8 I,10 I,9 I,10 >7 - -
Anal fin (A) 10 >8 10 >7 I,8 I,8 I,8 I,9 - - -
C (branched, segmented) 12 13 ? 13 12 12 12 13 - - -
C (unbranched, segmented)             2 + 1 or 2 2 + 2      
C (unbranched, unsegmented)           7 10 + ? 10 + 12      
Ventral fin (V) - - - - I,5 - I,5 I,5 - - -
Pectoral fin (P) - >11 14 >9 12 - >14 16 >11 >13 >9
Vertebrae [postabdominal vert.] >23 [17] >21 [16] >27 >23 [17] 29 [17] 30 [18] 30 [18] 31
[19]
>23 >11 29 [17]
Pterygiophore formula D1 - - - - - - - 3-2121100 - - -

 

 

TABLE 2. Morphometrics of the genus †Paralates in mm. Values standardized to standard length are given in brackets. Abbreviations: A = anal fin, C = caudal fin, D1 = first dorsal fin, D2 = second dorsal fin, D2C = distance between end of D2 and first procurrent ray of C, P = pectoral fin, V = pelvic fin.

# Ruf. 11/13_2 Ruf. 15a/b Ruf. 18 P 59784 P 59785 P 59786
Total length 29.1 (115%) 29.5 (118%) 34.5 (119%) 35.3 (116%) 22.5 (116%) >22.3
Standard length 25.2 25.1 29.1 30.4 19.4 >22.3
Max body height 5.4 (21.4%) 5.4 (21.5%) 5.5 (18.9%) 5.9 (19.4%) 4.4 (22.7%) -
Head length 9.4 (37.3%) 6.9 (27.5%) 7.1 (24.4%) 9.5 (31.3%) 5.6 (28.9%) 6.6
Head height 6.6 (26.1%) 5.4 (21.5%) 5.2 (17.9%) 6.9 (22.7%) 4.9 (25.3%) -
Horizontal eye diameter 1.5 (5.9%) 1.2 (4.8%) 1.3 (4.5%) - 1.7 (8.8%) 1.4
Dist snout to D1 11.7 (46.4%) 9.2 (36.7%) 10.8 (37.1%) 11.7 (38.5%) 7.8 (40.2%) 7.9
Dist snout to D2 14.6 (57.9%) 14.5 (57.8%) 16.2 (55.7%) 17.4 (57.2%) 11.2 (57.7%) 12.8
Dist snout to A 15.8 (62.7%) 15.7 (62.5%) 16.5 (56.7%) 18.1 (59.5%) 11.5 (59,3%) 13.3
Dist snout to P 9.4 (37,3%) 8.7 (34.7%) - 10.7 (35.2%) 6.8 (35.1%) 7.4
Dist snout to V 9.2 (36.5%) 8.2 (32.7%) - 10.6 (34.9%) 6.6 (34.0%) -
Length D1 >2.1 >2.4 >2.4 >3.4 >1.7 -
Length D2 2.8 (11.1%) >2.4 4.4 (15.1%) >4.3 >3.1 -
Length A 2.6 (10.3%) >2.4 >2.3 >3.5 >2.9 -
Length P >0.8 >1.3 - >1.9 >0.8 >2.8
Length V >1.2 >1.5 - >2.7 2.7 (13.9%) -
Base length D1 >0.6 2.3 (9.2%) 2.3 (7.9%) 2.6 (8.6%) 1.6 (8.2%) -
Base length D2 3.1 (12.3%) 3.0 (12.0%) 4.0 (13.7%) >3.5 2.4 (12.4%) -
Base length A 3.0 (11.9%) 2.5 (19.0%) 3.7 (12.7%) 3.6 (11.8%) 1.9 (9.8%) -
Length caudal peduncle 6.3 (25.0%) 6.8 (27.1%) 8.9 (30.6%) 8.3 (27.3%) 5.7 (29.4%) -
D2C 6.7 (26.6%) 7.3 (29.1%) 8.4 (28.6%) 7.5 (24.7%) 5.0 (25.8%) -
Min height of caud ped 2.0 (7.9%) 1.7 (6.8%) 2.6 (8.9%) 3.0( 9.7%) 2.2 (11.3%) -
Length of caudal fin 3.8 (15.1%) >3.8 >5.0 5.4 (17.8%) 3.6 (18.6%) -
Length of vertebrae 0.5-0.6 (1.9-2.3%) 0.6-0.7 (2.4-2.8%) 0.6-0.8 (2.1-2.7%) 0.5-0.7 (1.6-2.3%) 0.4-0.5 (2.1-2.6%) 0.4-0.5

 

TABLE 3. Meristic characters of recent gobioid taxa for which the pterygiophore formula 3-212110(0) has been observed in at least one species of the genus. The number of caudal fin rays refers to all segmented rays (branched and unbranched). Abbreviations: A = anal fin, C = caudal fin, D1 = first dorsal fin, D2 = second dorsal fin, P = pectoral fin, V = pelvic fin.

Taxon D1 D2 A C V P Vertebrae [postabd] Source Family
Paralates bleicheri Sauvage, 1883 VII I,10 I,9 >12-13 I,5 14 29
[17-18?]
this study inc. sed.
Paralates chapelcorneri n. sp. VII I,10 I,8-9 14-17 I,5 14-16 30?-31 [18?-19] this study inc. sed.
Gobiodon okinawae Sawada, Arai & Abe, 1972 VI I,10 I,9 15 - - 27
[17]
unpublished data Gobiidae
Gobiosoma hemigymnum (Eigenmann and Eigenmann 1888) VII I,11 I,9 17 I,5 17-19 27
[16]
van Tassell et al. 2015 Gobiidae
Gobiosoma spp. - - - - - - 27-28
[15-17]
Birdsong et al. 1988 Gobiidae
Gobulus spp. - - - - - - 27-28
[16-17]
Birdsong et al. 1988 Gobiidae
Gobulus spp. VII-VIII I,9-13 I,7-13 15-17 I,5 15-18 27-28
[16-17]
Hoese and Reader 2001 Gobiidae
Gobiomorphus alpinus Stokell, 1962 III-VII I,7-9 I,8-10 - - 16-19 29-31 McDowall 1994 Eleotridae
Gobiomorphus spp. III-VIII I,7-10 I,7-10 - - 13-21 27-31 McDowall 1975 Eleotridae
Gobiomorphus spp. - - - - - - 28-31
[16-19]
Birdsong et al. 1988 Eleotridae
Thalasseleotris       16     27
[17]
Hoese and Larson 1987 Thalasseleotrididae