TABLE 1. Species of Plesiosminthus described from the European Oligocene and Miocene.

Species Type-locality
P. schaubi Viret, 1926 (type-species) Coderet, France
P. myarion Schaub, 1930 Chavroches, France
P. promyarion Schaub, 1930 Puy-de-Montdoury, F rance
P. bavaricus Freudenberg, 1941 Gaimersheim, Germany
P. winistoerferi Engesser, 1987 Brochene Fluh 53, Switzerland
P. conjunctus Ziegler, 1994 Herrlingen 8, Germany
P. moralesi Álvarez Sierra, Daams and Lacomba Andueza, 1996 Sayatón 1, Spain
P. admyarion Comte, 2000 Thézels, France
P. meridionalis Comte, 2000 Venelles inf., France
P. margaritae n. sp. (this paper) Mirambueno 1, Spain
P. moniqueae n. sp. (this paper) Pech Desse, France
Plesiosminthus n. sp., Ziegler, 1994 Herrlingen 8, Germany
Plesiosminthus sp., Engesser and Mödden, 1987 Mümliswil-Hardberg, Switzerland
Plesiosminthus sp., Comte, 2000 Venelles inf., France

 

TABLE 2. Locality codes, species contents and most important reference of the localities studied. Zones are European mammal zones (Biochrom’97, 1997).

Code Locality Zone Species Reference
BF53 Brochene Fluh 53 MP30 P. winistoerferi Engesser (1987)
CANAL Canales MP28 Plesiosminthus sp. Alvarez Sierra et al. (1999)
CHAVR Chavroches MN2 P. myarion Hugueney and Vianey-Liaud (1980)
CLUZEL Cluzel MN2 P. myarion Hugueney and Vianey-Liaud (1980)
COD3 Coderet 3 MP30 P. schaubi Hugueney (1969)
DIEU Dieupentale MP30 P. schaubi Baudelot and Olivier (1978)
FORN11 Fornant 11 MN1 P. myarion Engesser (1987)
GAIM Gaimersheim MP28 P. bavaricus Kristkoiz (1992)
HERR8 Herrlingen 8 MP28 P. conjunctus Ziegler (1994)
LAUT2 Lautern 2 MN1 P. myarion Ziegler and Werner (1994)
MIR1 Mirambueno 1 MP27 P. margaritae this paper
MIR2A Mirambueno 2A MP27 P. aff. conjunctus this paper
OBL Oberleichtersbach MP30 P. winistoerferi Engesser and Storch (2008)
PARR Parrales MP29 P. schaubi Alvarez Sierra et al. (1999)
PDES Pech Desse MP28 P. promyarion Hugueney and Vianey-Liaud (1980)
PDES Pech Desse MP28 P. moniqueae this paper
PDF Pech-du-Fraysse MP28 P. promyarion Hugueney and Vianey-Liaud (1980)
RDUBEY Ruisseau du Bey MP28 P. promyarion Engesser (1987)
SAULC Saulcet MN1 P. myarion Hugueney and Vianey-Liaud (1980)
SAY1 Sayatón 1 MP29 P. moralesi Alvarez et al. (1996)
THZ Thézels MP30 P. admyarion Comte (2000)
VENINF Venelles inf. MP30 P. meridionalis Comte (2000)
VIV1 Vivel del Río 1 MP28 P. cf. margaritae this paper

 

TABLE 3. Frequencies of character states of the protolophule of M2, number of observations, and morphology values. Ant. = anterior, trans = transverse, ant.plus = anterior plus an incomplete posterior connection. * protolophule posterior, not double.

Sample ant. or trans ant. plus double n MV
CHAVR 0 3 97 33 0.985
FORN11          
VENINF 4 0 96* 25 0.960
THZ 95 0 5 97 0.050
COD3 97 0 3 63 0.032
HERR8 17 8 75 12 0.790
PDF          
GAIM 39 52 9 23 0.350
PDES 74 13 13 16 0.195
RDUBEY          
VIV1 57 4 39 69 0.410
MIR2A 19 7 74 27 0.775
MIR1 28 37 35 43 0.535

 

TABLE 4. P-values of t-test comparing the samples from THZ and COD3; df = degrees of freedom.

Molar Comte
(2000)
this paper df
m1 3.18 3.092 205
m2 2.02 -2.162 180
m3 3.00 2.828 127
M1 2.39 -2.483 188
M2 2.06 -2.178 172
M3 3.00 2.832 98

 

TABLE 5. Frequencies of character states of the protoconid hind arm of m2, number of observations, and morphology values. Data from Kristkoiz (1992, figure 84) and this paper.

Sample complete incomplete absent n MV
CHAVR 12 50 38 50 0.630
FORN11 26 52 22 27 0.480
VENINF 0 0 100 26 1.000
COD3 0 0 100 59 1.000
THZ 9 9 82 127 0.865
HERR8 82 18 0 17 0.090
PDF 36 53 11 28 0.375
GAIM 50 41 9 31 0.295
PDES 32 45 23 53 0.455
RDUBEY 31 69 0 13 0.345
VIV1 31 16 53 52 0.610
MIR2A 83 0 17 16 0.170
MIR1 60 26 14 41 0.270

 

TABLE 6. Frequencies of character states of the protoconid hind arm of m3, number of observations, and morphology values.

Sample complete incomplete absent n MV
CHAVR 5.4 0.0 94.6 37 0.946
VENINF 0.0 0.0 100.0 8 1.000
COD3 0.0 0.0 100.0 32 1.000
THZ 0.0 0.0 100.0 97 1.000
HERR8 0.0 0.0 100.0 14 1.000
PDF 8.0 32.0 60.0 25 0.760
GAIM 14.9 27.3 63.6 11 0.730
PDES 25.0 43.8 31.2 16 0.531
RDUBEY 0.0 0.0 100.0 6 1.000
VIV1 5.1 0.0 94.9 39 0.949
MIR2A 20.0 15.0 65.0 20 0.725
MIR1 23.2 11.6 65.1 43 0.710
 

TABLE 7. Number of specimens for samples in Figure 13.

Sample m1 m2 m3 M1 M2 M3
BF53 4 4 5 6 5 5
CHAVR 36 48 39 49 33 15
COD3 65 59 32 71 63 13
GAIM 34 32 11 28 23 3
HERR8 8 15 14 18 12 11
MIR1 17 41 45 49 43 37
PDES 57 54 16 35 16 3
SAY1 7 6 1 3 3 1
THZ 142 123 97 119 111 87
VENINF 19 25 8 27 26 1