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New and little known crane-fly species of the genera Helius, Elephantomyia and Toxorhina (Diptera, Limoniidae) from Dominican and Mexican amber

Katarzyna Kopeć, Iwona Kania, and Wiesław Krzemiński

Plain Language Abstract

Three new fossil crane-flies species—Helius (Helius) neali n. sp., Helius (Helius) oosterbroeki n. sp. and Toxorhina (Ceratocheilus) mexicana n. sp. from the Miocene fossil resins are described. The new representatives of Helius are characterized by, comparatively for these crane flies, a short rostrum, shorter or a little longer than the head. Additional information of known Miocene representatives of Elephantomyia are provided.

Resumen en Español

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Traducción: Enrique Peñalver (Sociedad Española de Paleontología) or Diana Elizabeth Fernández

Résumé en Français

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Translator: Kenny J. Travouillon or Antoine Souron

Deutsche Zusammenfassung

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Translator: Eva Gebauer

Arabic

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Translator: Ashraf M.T. Elewa

 

 

FIGURE 1. Elephantomyia (Elephantomyia) grata Podenas and Poinar, 2001, No. 5546 (sex unknown). 1, a drawing of the head (reconstructed) in latero-dorsal view; 2, a schematic representation of the relation between the length of palpi (p), antenna (a), rostrum (r) and head (h); 3, a drawing of the left wing venation; and 4, a drawing of the right antenna in latero-dorsal view.

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FIGURE 2. Elephantomyia (Elephantomyia) grata Podenas and Poinar, 2001, No. 5546 (sex unknown). 1, a photograph of the body in lateral view; and 2, a photograph of the right antenna in latero-dorsal view.

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FIGURE 3. Helius (Helius) neali n. sp., No. 10938 (male). 1, a drawing of the head (reconstructed) in fronto-dorsal view; 2, a schematic representation of the relation between the length of antenna (a), rostrum (r) and head (h); 3, a drawing of the left antenna in ventral view; 4, a drawing of the left wing venation; and 5, a drawing of the right gonocoxite and gonostyles in ventral view. Abbreviation: gx, gonocoxite; ing, inner gonostylus; oug, outer gonostylus.

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FIGURE 4. Helius (Helius) neali n. sp., No. 10938 (male). 1, a photograph of the body in fronto-dorsal view; and 2, a photograph of the head in fronto-dorsal view and the part of thorax in dorsal view.

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FIGURE 5. Helius (Helius) oosterbroeki n. sp., No. 10189 (male). 1, a drawing of the head (reconstructed) in fronto-ventral view; 2, a drawing of the right antenna in fronto-dorsal view; 3, a drawing of the right palpus in fronto-ventral view; 4, a schematic representation of the relation between the length of palpi (p), antenna (a), rostrum (r) and head (h); 5, a drawing of right gonocoxite and gonostyles in ventral view; and 6, a drawing of the right wing. Abbreviation: gx, gonocoxite; ing, inner gonostylus; oug, outer gonostylus.

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FIGURE 6. Helius (Helius) oosterbroeki n. sp., No. 10189 (male). 1, a photograph of the body in ventral view; 2, a photograph of the head in fronto-ventral view; and 3, a photograph of hypopygium in ventral view.

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FIGURE 7. Toxorhina (Ceratocheilus) mexicana n. sp., No. PI II 1870 (female). 1, a drawing of the head (reconstructed) in lateral view; 2, a schematic representation of the relation between the length of palpi (p), antenna (a), rostrum (r) and head (h); 3, a drawing of the left wing venation; and 4, a drawing of the ovipositor in latero-ventral view.

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FIGURE 8. Toxorhina (Ceratocheilus) mexicana n. sp., No. PI II 1870 (female). 1, a photograph of the body in lateral view; 2, a photograph of the apical part of rostrum in lateral view; and 3, a photograph of the ovipositor in latero-ventral view.

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TABLE 1. A list of fossil species of Limoniidae known from Dominican and Mexican amber.

Genus Species Locality
Chioneinae
Erioptera Erioptera (Mesocyphona) beata Podenas and Poinar, 2012 Mexico
Erioptera (Mesocyphona) divera Podenas and Poinar, 2012 Mexico
Erioptera (Mesocyphona) sherry Podenas and Poinar, 2012 Mexico
Gnophomyia Gnophomyia ieva Podenas and Poinar, 1999 Dominican Republic
Gnophomyia martini Podenas and Poinar, 1999 Dominican Republic
Molophilus Molophilus (Molophilus) theischingeri Krzemiński, 1992 Dominican Republic
Styringomyia Styringomyia dominicana Podenas and Poinar, 1999 Dominican Republic
Styringomyia optiova Podenas and Poinar, 2001 Dominican Republic
Limnophilinae
Epiphragma Epiphragma aurora Podenas and Poinar 2001 Dominican Republic
Polyneura Polyneura specula Podenas and Poinar, 1999 Dominican Republic
Polyneura virgo Podenas and Poinar, 1999 Dominican Republic
Limoniinae
Dicranomyia Dicranomyia (Caenolimonia) alexbrowni Podenas and Poinar, 2012 Mexico
Dicranomyia (Dicranomyia) chiapa Podenas and Poinar, 2012 Mexico
Dicranomyia (Dicranomyia) fera Podenas and Poinar, 1999 Dominican Republic
Dicranomyia (Dicranomyia) lema Podenas and Poinar, 1999 Dominican Republic
Dicranomyia (Dicranomyia) mexa Podenas and Poinar, 2012 Mexico
Dicranomyia (Dicranomyia) vela Podenas and Poinar, 2012 Mexico
Elephantomyia Elephantomyia grata Podenas and Poinar, 2001 Mexico
Geranomyia Geranomyia euchara Podenas and Poinar, 2001 Dominican Republic
Helius Helius collemus Podenas and Poinar, 2012 Mexico
Rhipidia Rhipidia mira Podenas and Poinar 1999 Dominican Republic
Trentepohlia Trentepohlia (Parangoma) agri Podenas and Poinar, 1999 Dominican Republic
Trentepohlia (Parangoma) immemorata Podenas and Poinar, 2001 Mexico
Trentepohlia (Parangoma) mexicana Podenas and Poinar, 2001 Mexico
 

author1Katarzyna Kopeć. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, 31-016 Kraków, Poland. This email address is being protected from spambots. You need JavaScript enabled to view it.

Katarzyna Kopeć received her Ph.D. in Biological Sciences at the Pedagogical University in Cracow in 2012. Since 2015 she works in the Natural History Museum of the Institute of Systematics and Evolution of Animals, Polish Academy of Sciences in Cracow. The area of her research covers the fossil and extant Mecoptera, and the Diptera.

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author2Iwona Kania. Department of Environmental Biology, University of Rzeszów, Zelwerowicza 4, 35-601 Rzeszów, Poland. This email address is being protected from spambots. You need JavaScript enabled to view it. - corresponding author

Iwona Kania, entomologist, palaeoentomologist, since 2007 Assistant Professor in Department of Environmental Biology, Faculty of Biology and Agriculture, University of Rzeszów. She graduated Ph.D. in 2007, in the Department of Zoology, University of Silesia in Katowice, and received habilitation degree in 2016. She authored and co-authored number of publications on taxonomic and phylogenetic issues of selected groups of insects, namely classification, morphology and biology of Diptera. Her scientific interest covers the palaeontology, systematics and evolution of recent and extinct Tipulomorpha flies. Her main research activities focus on inclusions of Tipulomorpha in fossil resins. She studied fossils from various collections in Poland and abroad, including these at Institute of Palaeontology, Russian Academy of Sciences in Moscow and Muséum National d'Histoire Naturelle in Paris. As Member of Board of International Palaeoentomological Society she attended Fossils X3 conferences in Beijing, China (2010) and Byblos, Lebanon (2013), Edinburgh (UK) (2016).

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author3Wiesław Krzemiński. Institute of Biology, Pedagogical University of Kraków, Podbrzezie 3, 31-054 Kraków, Poland. This email address is being protected from spambots. You need JavaScript enabled to view it.

Wiesław Krzemiński, entomologist, palaeontologist working on systematics, evolution and phylogeny of Diptera and Mecoptera, based on extinct and extant species. Since 1990 the Head of the Natural History Museum of the Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Kraków, Poland. A state professor since 2003. Academic position at the University of Rzeszów (2005–2009) and at the Pedagogical University in Kraków since 2010. Author of more than 120 publications; a keen participant of scientific expeditions to Antarctica, Venezuela, Columbia, Peru, Ecuador, Korea, Vietnam, China, Australia, New Zealand, Brazil. Organizer of over seventy exhibitions in several museums, mainly on fossil and living fauna and on nature protection. The founding member of the International Palaeoentomological Society (2001, Kraków); took part in all its following conferences in Pretoria, South Africa (2005), Alava, Spain (2007), Beijing, China (2010), Byblos, Lebanon (2013).