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New dragonflies and damselflies from Middle Miocene deposits in SW Bulgaria (Insecta: Odonata)

Andre Nel, Nikolay Simov, Vladimir Bozukov, and Milen Marinov

Plain Language Abstract

The first fossil dragonflies discovered in Bulgaria are described and illustrated. They belong to genera, which were present in both Western Europe and Siberia in the Neogene. Their fossil discoveries in Bulgaria provide a link between these two regions, which are now disconnected.

Resumen en Español

Nuevas libélulas y caballitos del diablo de depósitos del Mioceno Medio del SO de Bulgaria (Insecta: Odonata)

Se describen e ilustran los primeros fósiles de Odonata descubiertos en Bulgaria, es decir, el Aeshnidae Oligaeschna bulgariensis sp. nov., el Sieblosiidae Stenolestes rhodopensis sp. nov., y el Dysagrionidae Primorilestes magnificus sp. nov. Estos géneros están presentes tanto en Europa Occidental como en Siberia para el Neógeno. El descubrimiento de estos fósiles en Bulgaria proporciona una continuidad entre estas dos regiones, que en la actualidad están desconectadas.

Palabras clave: Aeshnidae; Sieblosiidae; Dysagrionidae; nuevas especies; Mioceno; Bulgaria

Traducción: Enrique Peñalver (Sociedad Española de Paleontología)

Résumé en Français

Nouvelles espèces de libellules et demoiselles (Insecta : Odonata) des dépôts du Miocène moyen du sud-ouest de la Bulgarie

Le premiers Odonata fossiles découverts en Bulgarie sont décrits et illustrés, i.e., l'aeshnidé Oligaeschna bulgariensis sp. nov., le sieblosiidé Stenolestes rhodopensis sp. nov., et le dysagrionidé Primorilestes magnificus sp. nov. Ces genres sont présents à la fois en Europe occidentale et en Sibérie pendant le Néogène. Ces découvertes de fossiles appartenant à ces genres en Bulgarie indiquent un lien entre ces deux régions qui sont maintenant déconnectées.

Mots-clés : Aeshnidae ; Sieblosiidae ; Dysagrionidae ; nouvelle espèce ; Miocène ; Bulgarie

Translator: Antoine Souron

Deutsche Zusammenfassung

Neue Libellen und Kleinlibellen aus dem mittleren Miozän von Südwest-Bulgarien (Insecta: Odonata)

Es werden die ersten fossilen Odonata aus Bulgarien beschrieben und illustriert, i.e., der Aeshnide Oligaeschna bulgariensis sp. nov., der Sieblosiide Stenolestes rhodopensis sp. nov. und der Dysagrionide Primorilestes magnificus sp. nov. Diese Gattungen kommen sowohl im Neogen Westeuropas als auch Sibiriens vor. Ihre Fossilfunde in Bulgarien verbinden diese beiden Regionen, die nun getrennt sind.

Schlüsselwörter: Aeshnidae; Sieblosiidae; Dysagrionidae; neue Art; Miozän; Bulgarien

Translator: Eva Gebauer

Arabic

Translator: Ashraf M.T. Elewa

 

 

FIGURE 1. Geographic position of the study site, Satovcha Graben, SW Bulgaria. Type locality of the new species is outlined with yellow.

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FIGURE 2. Oligaeschna bulgariensis sp. nov. 1, holotype Caт-1190a (part); 2, holotype Caт-1190b (part); 3, paratype Сат-55. Scale bars represent 5 mm in all figures.

 figure2

FIGURE 3. Stenolestes rhodopensis sp. nov. 1, holotype Caт-50a (part); 2, holotype Caт-50b (part); 3, 4, Caт-50a and Caт-50b - both wings; 5, paratype Caт-119 (whole insect); 6, paratype Caт-119 (wings only). Scale bars represent 5 mm in 1, 2, 3, 4, 6 ; 1 cm in 5.

figure3 

FIGURE 4. Primorilestes magnificus sp. nov. 1, holotype Сат-52 (part); 2, holotype Caт-53 (part); 3, paratype Caт-54. Scale bars represent 5 mm in all figures.

figure4

 

 

nelAndre Nel. Institut de Systématique, Évolution, Biodiversité, ISYEB - UMR 7205 - CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France. This email address is being protected from spambots. You need JavaScript enabled to view it.

André Nel obtained his PhD in 1991 from the University of Reims (France) working on the use of fossil insects in systematics, phylogeny and reconstructions of palaeoenvironments. As Maître de Conférence and Professor at the MNHN since 1993, he developed his studies on fossil insects on numerous orders, but especially on Odonata. He published ca. 550 papers on the subjects between 1983 and 2016.

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simovNikolay Simov. National Museum of Natural History, 1 Tsar Osvoboditel Blvd., 1000 Sofia, Bulgaria. This email address is being protected from spambots. You need JavaScript enabled to view it.

Nikolay Simov is a curator of Hemiptera at National Museum of Natural History, Sofia, Bulgaria. He works primarily on the biodiversity and taxonomy of Heteroptera, in particular those from Balkan Peninsula. His research interests also include paleontology, systematics, taxonomy, speleology and species conservation of dragonflies.

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bozukovVladimir Bozukov. Institute of Biodiversity and Ecosystem Research - Bulgarian Academy of Sciences, Acad. G. Bonchev Str., 23, BG-1113 Sofia, Bulgaria. This email address is being protected from spambots. You need JavaScript enabled to view it.

Vladimir Bozukov is an assistant professor at Department of Paleobotany and Palynology of IBER-BAS. He works on the morphology and taxonomy of Tertiary plants established by macrofossils. His research interests also include paleoecology, paleoclimatology and stratigraphy. He published ca. 50 papers on the subjects between 1995 and 2016.

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marinovMilen Marinov. Plant Health & Environment Laboratory, Investigation and Diagnostic Centre and Response, Ministry for Primary Industries, 231 Morrin Rd, Auckland, New Zealand. This email address is being protected from spambots. You need JavaScript enabled to view it.

Milen Marinov obtained his PhD in 2003 from the Institute of Zoology, Bulgarian Academy of Sciences. At present he is entomologist at the Ministry for Primary Industries, New Zealand. He works on the taxonomy, fauna and biogeography of the Pacific Odonata.