The fourteen case studies have been designed to demonstrate both the use of different data analysis methods in paleontology and the specific use of the functions in the program. The cases are taken from such diverse fields as morphology, taxonomy, paleoecology, paleoclimatology, sedimentology, extinction studies, and biostratigraphy. The examples are taken from both vertebrate and invertebrate paleontology, and they cover the whole of the Phanerozoic. These case studies are well suited for an introductory course in paleontological data analysis and have been tested in classroom situations. The cases are organized into four main subject areas: morphology and taxonomy, biogeography and paleoecology, time-series analysis, and biostratigraphy.
Case studies 1-5 involve the description and analysis of morphological
variation of different sorts, while case study 6 targets some phylogenetic
problems in a group of Cambrian trilobites and the mammals.
PE Note: The Case Study files are available from the PE
site, and also directly from the author. The links below point to the
author's site, which will, as time and the author proceed, contain updates and
newer versions.
Case
Study 1 investigates the external morphology of the Permian brachiopod Dielasma,
developing ontogenic models for the genus and comparing the growth rates and
outlines of different samples from in and around a Permian reef complex. In
a more focused exercise, Case
Study 2 uses spatial statistics to assess the mode of distribution of tubercles
on the cranidium of the trilobite Paradoxides from the middle Cambrian.
Case
Study 3 tackles the multivariate morphometrics of the Ordovician illaenid
trilobite Stenopareia using Principal Components Analysis (PCA), Principal
Coordinate Analysis (PCO), cluster and discriminant analyses to determine the
validity of two species from Scandinavia.
Case
Study 4 demonstrates the use of Elliptic Fourier shape analysis and principal
components for detecting changes in trilobite cephalon shape through ontogeny.
In
Case Study
5, aspects of the allometric growth of the Triassic rhynchosaur Scaphonyx
are investigated using regression analysis.
Case
Study 6 investigates the phylogenetic structure of the middle Cambrian Paradoxididae
through cladistic analysis, using parsimony analysis and bootstrapping. Similar
techniques can be applied to a matrix of 20 taxa of mammal; cladograms generated
by the program can be compared with a cluster analysis of the data matrix.
Case
studies 7-11 cover aspects of paleobiogeography and paleoecology. Case
Study 7 analyzes a global dataset of late Ordovician brachiopod distributions.
A series of provincial faunas were developed against a background of regression
and cooler surface waters during the first strike of the late Ordovician (Hirnantian)
glaciation. Through the calculation of similarity and distance coefficients
together with cluster analysis, these data can be organized into a set of latitudinally
controlled provinces. Seriation helps to develop any faunal, possibly climatically
generated, gradients within the data structure.
In
Case Study
8 faunal changes through a well-documented section in the upper Llanvirn
rocks of central Wales are investigated graphically and by the calculation of
diversity, dominance, and related parameters for each of ten horizons in the
sections. The changes in faunas fingerprint environmental shifts through the
section, shadowed by marked changes in lithofacies. This dataset is ripe for
considerable experimentation.
Case
Study 9 involves a re-evaluation of Ziegler's classic Lower Paleozoic depth-related
communities from the Anglo-Welsh area. Using a range of multivariate techniques
(similarity and distance coefficients, cluster analysis, detrended correspondence
analysis, and seriation) the reality and mutual relationships of these benthic
associations can be tested using a modified dataset.
Case
Study 10 discusses some well-known Jurassic shelly faunas from England and
France. The integrity and onshore – offshore distribution of six Corallian bivalve-dominated
communities is investigated with diversity measures, cluster analysis and detrended
correspondence analysis.
Case
Study 11 completes the analysis of biotic assemblages with an investigation
of the direction and orientation of a bedding-plane sample of brachiopod shells
from the upper Ordovician rocks of Scotland.
Two
cases involve the study of time series data. Case
Study 12 investigates the periodicity of mass extinctions during the Permian
to Recent time interval using spectral analysis. A number of diversity curves
can be modeled for the Paleozoic and post-Paleozoic datasets available in Fossil
Record 2, and turnover rates can be viewed for Phanerozoic biotas.
Case
Study 13 addresses the periodicity of oxygen isotope data from ice cores
representing the last million years of Earth history.
The
final case study (Case
Study 14) demonstrates the use of quantitative biostratigraphical correlation
with the method of Unitary Associations. Eleven sections from the Eocene of
Slovenia are correlated using alveolinid foraminiferans studied by Drobne.