Figure 3. Landmarks used to represent tooth crown morphology. A. Crown of the first lower right molar of Sorex araneus in functional occlusal view, in which the tooth is oriented so that the line of sight is parallel to all of the vertical shearing blades. The positions of the nine landmarks are indicated by open orange circles. B. Diagrammatic representation of the nine landmarks as portrayed in many of the simulation results. The trigonid and talonid landmarks are each connected by a thick grey line. The five cusp landmarks are labelled.