Etymology. The species named in honor of the outstanding investigator of Iranian geology Professor J. Stöcklin.
Holotype. GIN 4777/128. Axial section; Anarak, Absheni Formation, Unit 4; Carboniferous, Pennsylvanian, lower Moscovian.
Material. 6 axial and 5 tangential sections.
Description. Shell minute, inflated fusiform to subellipsoidal, with bluntly rounded poles. Mature specimens possess 4 to 6 volutions and measure from 0.9 to 1.25 mm in length and from 0.63 to 0.8 mm in diameter; form ratio 1.43-1.56. Early 1-1.5 volutions subdiscoidal to spherical and occasionally coiled at large angles to the coiling axis of following volutions. Coiling uniform but rather loose. Spirotheca composed of tectum, lower tectorium and indistinct upper tectorium 0.015 mm thick in outermost volution. Septa are thin. Septal fluting poorly developed in polar regions. Proloculus spherical, its outer diameter 0.07 mm. Tunnel rather narrow, about half as high as chambers. Chomata massive and wide.
Discussion. The species described closely resembles Profusulinella parva (Lee et al. 1930) but differs from the latter in having a more elongate shell and a more massive chomata. It is similar to P. copiosa (Thompson 1948) but differs from the latter in having a narrower tunnel and a more massive chomata.
Stratigraphic range. Carboniferous, Pennsylvanian, lower Moscovian.
Occurrence. Iran, Zaladu section, Absheni Formation, Unit 5, samples Z209, Z249; Anarak section, Absheni Formation, Unit 4, sample A120.
Etymology. The species named in honor of the outstanding investigator of Iranian geology Professor J. Stöcklin.
Holotype. GIN 4777/147. Axial section; Iran, Zaladu section, Absheni Formation, Unit 5; Carboniferous, Pennsylvanian, lower Moscovian.
Material. 8 axial and subaxial sections.
Description. Shell moderate in size, fusiform to inflate fusiform with straight to slightly convex lateral slopes and bluntly pointed poles. Mature specimens of 4.5 to 5 volutions and measure 3.0 to 3.6 mm in length and 1.3 to 1.6 mm in diameter; form ratio 2.15 to 2.4. Spirotheca composed of tectum and protheca (lower tectorium); poorly developed upper tectorium occurs only in the two early volutions. Septa thin, nearly plain, except for slight folding near the poles. Proloculus moderate in size; its outer diameter 0.10 to 0.17 mm. Tunnel moderately wide, except the last volution where it is very wide, about half as high as corresponding chambers. Chomata narrow, rounded, about half as high of chambers.
Discussion. Aljutovella stocklini most closely resembles Al. cafirniganica (Bensh 1969) but differs from the latter in fewer, but larger volutions, more rounded poles, and a less well developed upper tectorium in the early volutions. Aljutovella stocklini also possess slightly convex lateral slopes, whereas the lateral slopes of Al. cafirniganica are straight or slightly concave.
Stratigraphic range. Carboniferous, Pennsylvanian, lower Moscovian.
Occurrence. Iran, Zaladu section, Absheni Formation, Unit 5, sample Z227.
Etymology. The species name is derived from Iran.
Holotype. GIN 4777/157. Axial section. Iran, Zaladu section, Absheni Formation, Unit 5; Carboniferous, Pennsylvanian, lower Moscovian.
Material. 4 axial sections.
Description. Shell moderate in size possesses inflated fusiform shape, with straight to slightly convex lateral slopes and bluntly pointed poles. Mature specimens of 5 to 6 volutions measured 2.5 to 3.1 mm in length and 1.45 to 1.75 mm in diameter; form ratio 1.65 to 1.7. Spirotheca composed of tectum and protheca; poorly developed upper tectorium occurs only in two early volutions. Septa thin, wavy across length of shell and gently folded near the poles. Proloculus of medium size; its outer diameter 0.08 to 0.10 mm. Tunnel narrow in the early volutions and wide in the outer ones. Chomata narrow and low.
Discussion. Aljutovella iranica is most like Aljutovella postaljutovica (Safonova in Rauser-Chernousova et al. 1951) but differs from the latter in having a less well- developed chomata, loosely coiled volutions, and a larger size of corresponding volutions. Aljutovella iranica differs from Al. cafirniganica (Bensh 1969) and Al. stocklini in having a more inflated and rounded shell, a less well-developed chomata, and more folded septa.
Stratigraphic range. Carboniferous, Pennsylvanian, lower Moscovian.
Occurrence. Iran, Zaladu section, Absheni Formation, Unit 5, samples Z219 and Z220.
Etymology. The species is named in honor of Dr. M.N. Gorgij, who studied the Zaladu section and collected fusulinid samples there.
Holotype. GIN 4777/157. Axial section. Iran, Zaladu section, Absheni Formation, Unit 5; Carboniferous, Pennsylvanian, lower Moscovian.
Material. 3 axial and 4 subaxial and tangential sections.
Description. Shell moderate in size, fusiform, with straight to slightly convex lateral slopes and bluntly pointed poles. Mature shells of 4.5 to 5 volutions are measured as 2.6 to 2.75 mm in length and 1.2 to 1.25 mm in diameter; form ratio of 2.1 to 2.25. Spirotheca are composed of tectum, lower and upper tectoria in the early volutions and tectum and protheca in the late ones. Septa are thin and nearly in one plane across the middle of the shell or gently folded near poles. Proloculus is relatively large, its outer diameter 0.08 to 0.125 mm. Tunnel is wide and low throughout. Chomata are small and rounded.
Discussion. The species described is greatly similar to the forms reported as Aljutovella ex gr. Al. distorta (Leontovich in Rauser-Chernousova et al. 1951) from the Bashkirian-Moscovian boundary beds of Asturias, Spain (Granados et al. 1985). It is distinguished from Al. distorta by having an inflated shell, larger shell dimensions, less pronounced chomata, and a wider tunnel. Aljutovella gorgiji differs from Al. iranica in having a more elongate shell and straighter septa in the middle part of the shell. Unlike Al. stocklini, Al. gorgiji is smaller, more elongate, has a less massive chomata, and wavier septa.
Stratigraphic range. Carboniferous, Pennsylvanian, lower Moscovian.
Occurrence. Iran, Zaladu section, Absheni Formation, Unit 5, sample Z220.
Material. 18 axial and subaxial sections.
Description. Shell fusiform to inflated fusiform, with slightly convex to slightly concave lateral slopes and bluntly pointed poles. Mature specimens of 4 to 5 volutions measured as 1.72 to 2.87 mm in length and 1.0 to 1.43 mm in diameter; form ratio 1.66 to 2.2. Volutions coiled loosely throughout beginning from the second volution. Spirotheca thin, its outermost volution reaches 0.02 to 0.03 mm. Septa is thin, nearly in one plane across the middle of shell, and gently folded near poles. Proloculus relatively large, its outer diameter 0.08 to 0.25 mm. Tunnel is moderately wide, about half as high as chambers. Chomata is narrow, about half as high as chambers.
Remark. Due to poor preservation, we hesitate to consider specimens to represent a new species. These specimens are distinguished from all known Aljutovella species, by a very loose, irregular spiral coiling and variable shape of the shell. They are most like Aljutovella subaljutovica (Rauser-Chernousova et al. 1951) but differ from the latter in having a more rounded shell, fewer volutions, and an irregular spiral coiling.
Stratigraphic range. Carboniferous, Pennsylvanian, lower Moscovian.
Occurrence. Iran, Zaladu section Absheni Formation, Unit 5, sample Z191.
Remarks. The single specimen described is close to Fusulinella (Moellerites) bockiformis (Bogush 1963), but a greater developed diaphanotheca allows it to be attributed to the subgenus Fusulinella (Fusulinella) (Möeller 1877). In addition, this specimen has a more inflated shell than Fusulinella (M.) bockiformis, which is nearly spherical in the early volutions. This fusilinid may represent a new species, however, limited material prevents us from naming it at this time.
Stratigraphic range. Carboniferous, Pennsylvanian, lower part of the upper Moscovian.
Occurrence. Iran, Anarak section, Absheni Formation, Unit 5, sample A184.
Etymology. The species name derived from ancient name of Iran – Persia.
Holotype. GIN 4777/39. Axial section; Iran, Anarak section, Absheni Formation, Unit 5; Carboniferous, Pennsylvanian, lowermost part of upper Moscovian.
Material. 14 axial sections.
Description. Shell large, inflated fusiform to elongate fusiform, with straight to slightly convex lateral slopes and bluntly pointed poles; first one or twovolutions often subrhomboidal. Mature specimens of 5 volutions measured as 3.5 to 5.5 mm in length and 1.4 to 2.0 mm in diameter; form ratio 2 to 2.94. Thick spirotheca composed of tectum and protheca; wall penetrated by very fine, but prominent pores; upper tectorium occasionally present in one or two early volutions. Thickness of the spirotheca in two late volutions varies from 0.04 to 0.06 mm. Thin septa intensely and relatively regularly fluted from pole to pole. Proloculus spherical; its outer diameter 0.1 to 0.17 mm. Tunnel narrow in the early volutions and wide in the ones that follow. Massive chomata present in two early volutions.
Discussion. Putrella persica closely resembles Putrella brazhnikovae fusiformis (Putrja 1948) but is distinguished from the latter by having more inflated early volutions, less regular septal fluting, and more massive and wider chomata in the early volutions. The most elongate specimens of Putrella persica sp. nov. resemble P. brazhnikovae brazhnikovae (Putrja 1948), but differ from the latter in a slower rate of shell lengthening throughout the shell, more regular coiling of volutions, and stronger, more regular septal fluting. The other difference is that Putrella persica possess three-layered spirotheca in the early volutions, in contrast to four-layered spirotheca with diaphanotheca in the early volutions of P. brazhnikovae. Putrella persica differs from P. admiranda (Leven 1998) by less regular septal fluting and more compact coiling of the early volutions.
Stratigraphic range. Carboniferous, Pennsylvanian, lower part of upper Moscovian.
Occurrence. Iran, Anarak section, Sardar 2 Sub-formation, Unit 5, samples A183 and A184.
Material. 1 axial section.
Description. Shell moderately large, inflate fusiform, with slightly convex lateral slopes and bluntly rounded poles. Mature shells have only 3.5 loosely coiled volutions and measure 4.5 mm in length and 1.85 mm in diameter; form ratio 2.4. Spirotheca composed of tectum and thick (0.4 mm) protheca. Septa thin and very irregularly fluted from pole to pole except for the tunnel area. Septal folds are high. Proloculus is very large, its outer diameter 0.06 mm. Tunnel low and wide throughout. Very small chomata present only on proloculus.
Discussion. Putrella sp. 1 differs from all known Putrella species in large proloculus and loosely coiled volutions. The specimen most probably represents a new species, but due to limited material we cannot diagnose it at this time.
Stratigraphic range. Carboniferous, Pennsylvanian, lower part of upper Moscovian.
Occurrence. Iran, Anarak section, Absheni Formation, Unit 5, sample A183.
Material. 1 axial section.
Description. Shell is large, inflated fusiform, with straight or slightly convex lateral slopes and bluntly pointed poles. Mature specimens have six volutions, the early four of which are tightly coiled; coiling of last two volutions looser. Spirotheca composed of tectum and thick protheca; thickness of spirotheca in outer volution 0.056 mm. Septa thin, irregularly fluted from pole to pole except for the tunnel area. Septal folds low to very low. Proloculus of moderate size; its outside diameter 0.125 mm. Tunnel very narrow and high in tightly coiled early volutions and wide and low in the loosely coiled ones.
Discussion. Putrella sp. 2 differs from other Putrella species in strongly irregular coiling of volutions and in relatively weak and low septal fluting. Similarly to the previous specimens, Putrella sp. 1, we cannot establish new species due to limited material.
Stratigraphic range. Carboniferous, Pennsylvanian, lower part of upper Moscovian.
Occurrence. Iran, Anarak section, Absheni Formation, Unit 5, sample A183.
Etymology. The species is named for the Anarak section where it was found.
Holotype. GIN 4777/184. Axial section; Iran, Anarak section, Absheni Formation, Unit 5; Carboniferous, Pennsylvanian, lowermost part of the upper Moscovian.
Material. 2 axial and 3 subaxial and tangential sections.
Description. Shell small, inflated fusiform to subrhomboidal, with straight to slightly concave lateral slopes and bluntly rounded poles. Mature shells possess 5.5 to 6 volutions and measure 2.3 to 3.1 mm in length and 1.25 to 1.65 mm in diameter; form ratio 1.87 to 2.0. Spirotheca composed of tectum, lower and upper tectoria in the early three volutions; distinct diaphanotheca appear at the fourth volution. Thickness of spirotheca in two outer volutions varies from 0.03 to 0.04 mm. Septa thin, moderately and irregularly fluted from pole to pole except for the tunnel area where septa weakly fluted or even straight. Proloculus is of moderate size; its outside diameter 0.14 to 0.2 mm. Tunnel narrow and high. Chomata high and massive.
Discussion. The weakly developed diaphanotheca of the Beedeina bona anarakensis n. subsp. suggests that this subspecies is a primitive Beedeina. The subspecies described resembles, in many respects, the nominative subspecies, but differs from the latter in having weaker septal fluting. This feature makes it similar to some forms of ancestral genus Citronites, such as C. citronoides (Manukalova 1948). However, the presence of pronounced diaphanotheca prevents assigning the species to this genus.
Stratigraphic range. Carboniferous, Pennsylvanian, lower part of upper Moscovian.
Occurrence. Iran, Anarak section, Absheni Formation, Unit 5, samples A184 and A193.
