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TABLE 1. Overview on some systematic assignments of castorid taxa by different authors on the level of genera only. In blue those genera that are assigned to the Palaeocastorinae by Korth (1994). Flynn and Jacobs (2008) refer to the North American Fauna only.

 

Stirton 1935

Simpson 1945

Korth 1994

McKenna & Bell 1997

Korth 2001

Rybczynski 2007

Flynn & Jacobs 2008

 

semi flattened incisors

Uncertain subfam.

Agnotocastorinae

 

basal castorids

 

Agnotocastor

 

Agnotocastor

 

Agnotocastorini

 

Agnotocastor

 

Palaeocastor

 

Hystricops

 

Agnotocastor

Agnotocastor

Neatocastor

 

Euhapsis

     

Neatocastor

Anchitheriomys

Oligotheriomys

 

Chalicomys

     

Hystricops

 

Anchitheriomys

 

Castor

     

Anchiteriomyini

     

Convex incisors

     

Anchitheriomys

     

Steneofiber

     

Propalaeocastor

     

Monosaulax

     

Oligotheriomys

     

Eucastor

Castorinae

Castorinae

Castorinae

Castorinae

Castorinae

Castorinae

 

Dipoides

Agnotocastor

Castor

Agnotocastor

Castor

Castor

Castor

 

Castoroides

Palaeocastor

 

Capacikalini

Chalicomys

Hystricops

   

Amblycastor

Euhapsis

 

Capacikala

Steneofiber

Sinocastor

   

Hystricops

Chalicomys

 

Pseudopalaeocastor

Zamolxifiber

Steneofiber

   

Trogontherium

Sinocastor

 

Castorini

Romanofiber

     

Conodontes

Castor

 

Euhapsina

Schreuderia

     
     

Euhapsis

       
     

Fossorcastor

       
     

Castorina

       
     

Propalaeocastor

       
     

Chalicomys

       
     

Palaeocastor

       
     

Hystricops

       
     

Castor

       
 

Castoroidinae

Castoroidinae

Castoroidinae

Castoroidinae

Castoroidinae

Castoroidinae

 
 

Steneofiber

Monosaulax

Steneofiber

Castoroidini

Dipoides

Euroxenomys

 
 

Monosaulax

Eucastor

Neatocastor

Monosaulax

Eucastor

Monosaulax

 
 

Eucastor

Dipoides

Asiacastor

Eucastor

Trogontherium

Eucastor

 
 

Dipoides

Procastoroides

Youngofiber

Dipoides

Euroxenomys

Microdipoides

 
 

Procastoroides

Castoroides

Trogontherium

Castoroides

Procastoroides

Dipoides

 
 

Castoroides

 

Eucastor

Procastoroides

Castoroides

Procastoroides

 
 

Trogontherium

 

Schreuderia

Trogontheriini

     
 

Amblycastor

 

Dipoides

Trogontherium

     
     

Boreofiber

Boreofiber

     
     

Romanofiber

Euroxenomys

     
     

Zamolxifiber

Youngofiber

     
     

Procastoroides

Asiacastor

     
     

Castoroides

       
     

Paradipoides

       
   

Palaeocastorinae

 

Palaeocastorinae

Palaeocastorinae

Palaeocastorinae

 
   

Palaeocastorini

 

Palaeocastor

Euhapsis

Palaeocastor

 
   

Palaeocastor

 

Capacikala

Fossorcastor

Capacikala

 
   

Capatanka

 

Pesudopalaeocastor

Pseudopalaeo-castor

Pseudopalaeocastor

 
   

Capacikalini

 

Euhapsis

Palaeocastor

Euhapsis

 
   

Capacikala

 

Fossorcastor

 

Fossorcastor

 
   

Pseudopalaeocastor

       
   

Euhapsini

         
   

Euhapsis

         
   

Fossorocastor

         
         

ungrouped

   
         

Migmacastor

   
         

Agnotocastor

   
         

Eutypomys

   
         

Priusaulax

   

Table 2. Measurements of the skull and mandible of Capacikalagradatus from the John Day fossil Beds in millimeters.

 

greatest skull length

59.93

zygomatic width (estimated from preserved right side)

48.00

length of right zygomatic arch

24.04

width of skull at external auditory meatus

30.43

distal height of skull (condyle - sagittal crest)

19.87

height of skull at auditory bullae

23.45

angle occipital plane to base of skull

90°

height of skull at tooth row (at P4)

22.97

width across condyles

12.52

maximal width of the nasals

9.25

maximal width of snout

12.39

height of snout

12.55

diastema

19.34

length of incisive foramen

5.24

a second foramen in front of incisive foramen present?

yes

form of infraorbital foramen

slitlike

height of infraorbital foramen

3.03

infraorbital constriction

8.66

width of parietale behind orbitae

22.62

width of postzygomatic constriction

25.05

length of right tooth row

11.25

length of left tooth row

11.34

distance P4/P4

4.31

distance M3/M3

7.2

postzygomatic skull length

14.58

length of bulla

11.46

tooth row visible trough orbital constriction

yes

rugosities on parietale

yes

masseter superficialis process prominent

no

masseter superficialis process clearly bordered

no

form of crista facialis in lateral view

straight/diagonally

form of crista facialis or rather infraorbital foramen in frontal view

convex, diagonally

lacrimal with dorsal part ?

no

Table 3. Measurements of the teeth of Capacikalagradatus JODA 621 in millimeters. Length of tooth rows: mandibular right 12.65 mm, mandibular left 12.72 mm, maxillar right 12.21 mm and maxillar left 11.27 mm.

 

Tooth

Length

Width

p4 left

4.27

3.33

m1 left

2.91

3.52

m2 left

2.92

3.44

m3 left

2.81

2.75

p4 right

4.13

3.32

m1 right

2.81

3.6

m2 right

3.08

3.47

m3 right

2.72

2.79

P4 left

3.43

3.81

M1 left

2.92

3.56

M2 left

2.65

3.44

M3 left

2.41

2.8

P4 right

3.57

4.11

M1 right

3.11

3.9

M2 right

2.89

3.56

M3 right

2.28

2.92

 

 

 

Table 4. Some values of procumbency angles of the upper incisors of some castorid taxa measured in accordance to the method of Lessa and Thaeler (1989) either from material at hand or the cited literature. Taxonomic assignments as in the cited literature. MTD – Senckenberg Natural History Collections Dresden, Museum für Tierkunde, UCMP – University of California Berkeley Museum of Paleontology.

 

Taxon

Literature

procumbency angle

Castoroides ohioensis

cast, MTD

63°

Capatanka magnus

Martin, 1987, fig. 2

69°

Monosaulax tedi

Korth, 1999, fig. 1

70°

Euhapsis breugerorum

Samuels & Van Valkenburgh, 2009

80°

Euhapsis platyceps

Samuels & Van Valkenburgh, 2009

82°

Castor fiber

average of 10 specimens, MTD

88°

Castor canadensis

Thomas, 1916

90°

Palaeocastor simplicidens

Samuels & Van Valkenburgh, 2009

90°

Capacikala gradatus

JODA 621, cast

95°

Palaeocastor fossor

Martin 1987, fig. 2

95°

Capatanka cankpeopi

McDonald, 1963

99°

Palaeocastor fossor

Samuels & Van Valkenburgh, 2009

102°

Dipoides tanneri

Korth, 2007, fig. 8

103°

Monosaulax pansus

Stefen, 2001

110°

Palaeocastor nebrascensis

UCMP 114635

113°

Palaeocastor magnus

Samuels & Van Valkenburgh, 2009

114°

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