TABLE 1. Experimental design and burrows produced. Sediment density is given in kgf/cm2. 1, Experiment 1; 2, Experiment 2; 3, Experiment 3. Sediment moisture values are in percent total volume. VS: vertical shaft, VC: vertical shaft with terminal chamber, SS: subvertical shaft, SC: subvertical shaft with terminal chamber, JS: J-shaped burrow, YS: Y-shaped burrow, IC: isolated chamber.

1. Experiment 1: Basic Morphology
Species Individuals Terrarium Size (gal) Sediment Composition Sediment Depth (cm) Sediment Density Sediment Moisture Duration (day) Resulting Burrow Architectures
G. inermis 5 20 30-50-20% organic-sand-clay/silt 35 0.35 55% 14 VS (3), SS (1), IC (1)
G. inermis 5 20 30-50-20% organic-sand-clay/silt 35 0.35 55% 30 VS (2), VC (2)
G. inermis 3 30 30-50-20% organic-sand-clay/silt 25 0.35 55% 14 VS (1), VC (1), SC (1)
G. inermis 3 30 30-50-20% organic-sand-clay/silt 25 0.35 55% 30 VS (1), SS (1)
H. lenta 5 10 50-50% organic-clay/silt 25 0.35 35% 14 VS (3), YS (1)
H. lenta 4 10 50-50% organic-clay/silt 25 0.35 35% 30 VS (2), VC (1)
H. lenta 4 20 50-50% organic-clay/silt 26 0.35 35% 14 VS (1)
                 
2. Experiment 2: Sediment Composition
Species Individuals Terrarium Size (gal) Sediment Composition Sediment Depth (cm) Sediment Density Sediment Moisture Duration (day) Resulting Burrow Architecture
G. inermis 5 20 30-35-35% organic-sand-clay/silt 35 0.35 55% 30 VS (4)
G. inermis 5 20 30-20-50% organic-sand-clay/silt 35 0.70 55% 30 VS (2), VC (2)
H. lenta 3 20 25-75% organic-clay/silt 26 0.35 35% 30 VS (2), SS (1)
H. lenta 4 20 100% clay/silt 26 0.70 35% 30 VS (1), SS (3)
                 
3. Experiment 3: Sediment Moisture
Species Individuals Terrarium Size (gal) Sediment Composition Sediment Depth (cm) Sediment Density Sediment Moisture Duration (day) Resulting Burrow Architecture
G. inermis 4 20 30-50-20% organic-sand-clay/silt 35 0.35 35% 30 VS (2), SS (1), JS (1)
G. inermis 4 20 30-50-20% organic-sand-clay/silt 35 0.70 75% 30 VS (1)
H. lenta 4 20 50-50% organic-clay/silt 26 0.35 15% 30 VS (2)
H. lenta 4 20 50-50% organic-clay/silt 26 0.35-0.70 55% 30 VS (1), SS (1)
                 

TABLE 2. Qualitative data for each burrow architecture. Burrows are arranged by increasing complexity. That distribution architectures produced by each species is provided, as well as the sum of all burrow architectures developed. (PDF only)

TABLE 3. Quantitative measurements of vertical shafts produced by Gorgyrella inermis and Hogna lenta. (PDF only)

TABLE 4. Quantitative measurements of subvertical shafts produced by Gorgyrella inermis and Hogna lenta.

  G5 G10 G12 G20 G24 H2 H11 H12 H13 H14 H16 H20
Surface Openings 1 1 1 1 1 1 1 1 1 1 1 1
                         
Maximum depth 11.2 13.9 6.6 4.4 8.3 5.5 5.5 6.1 4.4 5.7 6.3 6.6
                         
Total Length 13.2 14.4 7.8 4.6 8.4 5.7 5.6 6.2 5.2 5.8 6.9 7.4
                         
Maximum width 3.1 2.4 3.3 2.9 3.4 2.8 2.9 2.7 2.5 3.1 2.6 3.1
Minumum width 2.3 1.5 2.1 1.8 1.6 2.1 1.8 1.6 1.2 2.3 1.7 1.7
Mean width 2.6 2 2.9 2.3 2.6 2.3 2.4 2.3 2.2 2.7 2.3 2.5
                         
Maximum height 3 2.4 3 3 2.7 2.8 2.6 2.8 2.5 2.7 2.6 3.6
Minimum height 2.1 1.8 1.9 1.5 1.8 1.9 1.6 1.6 1.4 2.3 1.2 1.9
Mean height 2.3 2.1 2.5 2.4 2.4 2.2 2.2 2.4 2.2 2.5 2.3 2.8
                         
Mean W/H ratio 1.1 1 1.1 1 1.1 1 0.9 1 1 0.9 1 1.1
                         
Maximum circumference 8.9 7.6 9.5 8.4 9.4 8.9 9 8.8 7.9 9.1 8.3 11.1
Minimum circumference 7.4 5.7 4.5 5.6 5.4 6.1 4.5 5.1 3 7.1 4.8 6
Mean circumference 7.9 6.7 8.4 7.3 7.8 7.3 7.5 7.4 6.5 8.3 7.2 8.7
                         
Maximum slope 75° 75° 70° 70° 70° 75° 60° 65° 70° 60° 60° 76°
Minimum slope 75° 75° 70° 70° 70° 75° 60° 65° 70° 60° 60° 65°
Mean slope 75° 75° 70° 70° 70° 75° 60° 65° 70° 60° 60° 70.5°
                         
Branching angles NA NA NA NA NA NA NA NA NA NA NA NA
                         
Complexity 2 2 2 2 2 2 2 2 2 2 2 2
                         
Tortuosity 1.1 1 1.1 1 1 1 1.1 1.1 1.1 1 1 1.1
                         

TABLE 5. Quantitative measurements of other burrow architectures produced by Gorgyrella inermis and Hogna lenta. See Table 1 for architecture key.

  G2 G4 G16 G17 G26 H6 G27 G6 G23 H4
Architecture VC VC VC VC VC VC SC IC JS YS
                     
Surface Openings 1 1 1 1 1 1 1 1 1 2
                     
Maximum depth 16.3 7.08 13 12.5 5.7 3.2 9.3 2.1 8.6 4.1
                     
Total Length 18.4 7.7 13.2 12.5 5.7 3.2 9.5 3.7 8.9 6.7
                     
Maximum width 2.4 2.2 6 4.4 5.7 2.6 5.8 2.7 2 2.1
Minumum width 1.9 1.7 1.8 2.3 2.9 2.4 2.4 2.1 1.7 1.7
Mean width 2.1 1.9 3.6 3 4.3 2.5 3.5 2.5 1.9 1.9
                     
Maximum height 2.7 1.7 3.7 2.7 3.4 2.8 3.9 2.3 2 1.9
Minimum height 1.7 1.4 2.2 1.8 2.1 2.8 2.3 1.6 1.2 1.4
Mean height 2.1 1.6 2.7 2.2 2.9 2.8 3.1 1.9 1.7 1.8
                     
Mean W/H ratio 1 1.2 1.3 1.3 1.5 1.1 1.1 1.3 1.1 1
                     
Maximum circumference 7.9 6.3 15.6 12.2 23.6 7.8 15.3 7.9 6.4 6.8
Minimum circumference 5.4 5.2 3.1 6.9 17 5.1 7.2 7 5.1 4.9
Mean circumference 6.6 5.6 10.7 8.4 20.8 6.8 10.8 7.4 5.7 6.0
                     
Maximum slope 90° 90° 90° 90° 90° 85° 75° 45° 90° 75°
Minimum slope 78° 76° 90° 90° 90° 85° 75° 45° 39° 35°
Mean slope 84° 83° 90° 90° 90° 85° 75° 45° 64.5° 55°
                     
Branching angles NA NA NA NA NA NA NA NA NA 90°
                     
Complexity 3 3 3 3 3 3 3 2 2 4
                     
Tortuosity 1.4 1.2 1 1 1 1 1 1.1 1.4 1
                     

TABLE 6. Mann-Whitney and Kolmogorov-Smirnov test results ( p values) from the comparison of the burrows produced by arthropods considered in this study. 1) Gorgyrella and Hogna, 2) Spiders and Aphonopelma, 3) Spiders and Hadrurus, 4) Spiders and Mastigoproctus, 5) Spiders and Narceus. Boxed values indicate statistical difference ( p <0.05). All measurements are in cm, and all angles are in degrees.

1. Gorgyrella inermis and Hogna lenta
Organism Test Depth Length Width Height W/H Ratio Circum Slope Complexity Tortuosity
Spiders MW 0.006505 0.007414 0.4677 0.4947 0.002494 0.6257 0.5671 0.3968 0.3662
KS 0.005944 0.0005882 0.08522 0.325 0.02798 0.317 0.6544 0.2579 0.4907
                     
      G. inermis H. lenta
Analysis of Differences   Property Mean Median SD Range Mean Median SD Range
    Depth 8.9 8.45 4.22 16.6-2.1 5.78 5.7 1.47 8.7-3.2
    Length 9.36 8.5 4.38 18.4-2.5 6.07 6.2 1.42 8.7-3.2
    W/H Ratio 1.10 1.1 0.124 1.5-1 1.01 1.0 0.0809 1.2-0.90
                     
2. Spiders and Aphonopelma chalcodes
Organism Test Depth Length Width Height W/H Ratio Circum Slope Complexity Tortuosity
A. chalcodes MW 0.6675 0.4484 0.2795 0.7426 0.01879 0.03599 0.01724 0.2757 0.02417
KS 0.6696 0.6128 0.02454 0.5568 0.01319 0.02991 0.01319 0.6128 0.20454
                     
      A. chalcodes G. inermis and H. lenta
Analysis of Differences   Property Mean Median SD Range Mean Median SD Range
    W/H Ratio 0.659 0.92 0.156 1.3-0.5 1.06 1.0 0.117 1.5-0.9
    Circum 14.3 8.43 6.85 9.46-7.4 7.86 7.4 2.31 20.8-5.3
    Slope 26.3 35 1.77 45-25 79.56 85 11.27 90-45
    Tortuosity 0.754 1.04 0.326 1.1-0.98 1.02 1.0 0.124 1.4-0.50
                     
3. Spiders and Hadrurus arizonensis
Organism Test Depth Length Width Height W/H Ratio Circum Slope Complexity Tortuosity
H. arizonensis MW 0.07281 4.14E-07 5.03E-09 9.35E-05 1.32E-09 1.43E-08 1.10E-09 2.04E-09 3.38E-08
KS 0.09865 1.04E-05 2.70E-10 0.0005116 3.23E-12 2.70E-10 1.01E-11 1.52E-08 2.13E-09
                     
      H. arizonensis G. inermis and H. lenta
Analysis of Differences   Property Mean Median SD Range Mean Median SD Range
    Total Length 26.9 23 18.5 76-7 8.03 6.9 3.84 18.4-2.5
    Width 4.34 4.4 0.553 5.2-3.1 2.48 2.3 0.515 4.3-1.7
    Height 1.86 1.9 0.347 2.4-1.0 2.33 2.3 0.391 3.2-1.6
    W/H Ratio 2.41 2.4 0.434 3.1-1.7 1.06 1.0 0.117 1.5-0.9
    Circum 12.3 12.3 1.96 17.3-9 7.86 7.4 2.31 20.8-53
    Slope 17.2 15 11.7 45-5 79.6 85 11.3 90-45
    Compexity 4.00 3 2.12 9-3 2.21 2 0.463 4-2
    Tortuosity 1.96 1.3 1.94 9.0-1.1 1.02 1.0 0.124 1.4-0.50
                     
4. Spiders and Mastigoproctus giganteus
Organism Test Depth Length Width Height W/H Ratio Circum Slope Complexity Tortuosity
M. giganteus MW 0.9532 0.001977 4.45E-05 0.01405 1.47E-09 1.58E-05 9.70E-06 0.01234 0.002202
KS 0.8078 0.005419 4.45E-05 0.01874 6.13E-10 2.10E-05 2.10E-05 0.07236 0.002164
                     
      M. giganteus G. inermis and H. lenta
Analysis of Differences   Property Mean Median SD Range Mean Median SD Range
    Length 17.7 14.25 15.4 66-3.8 8.03 6.9 3.84 18.4-2.5
    Width 3.367 3.4 1.13 5.8-2.1 2.48 2.3 0.515 4.3-1.7
    Height 2 1.9 0.512 2.9-1.0 2.33 2.3 0.391 3.2-1.6
    W/H Ratio 1.82 1.9 0.342 2.4-1.2 1.06 1.0 0.117 1.5-0.9
    Circum 11.0 10.65 2.30 17-7.3 7.86 7.4 2.31 20.8-53
    Slope 55.4 50 21.1 90-20 79.6 85 11.3 90-45
    Compexity 3.17 2 2.12 10-2 2.21 2 0.463 4-2
    Tortuosity 1.31 1.25 0.363 2.4-1.0 1.02 1.0 0.124 1.4-0.50
                     
5. Spiders and Narceus americanus
Organism Test Depth Length Width Height W/H Ratio Circum Slope Complexity Tortuosity
N. americanus MW 1.01E-05 0.1036 6.60E-09 6.80E-09 0.2743 7.09E-09 0.007136 0.444 0.0006592
KS 1.17E-05 0.05088 3.24E-11 1.00E-04 0.05088 3.24E-11 0.04083 0.004083 0.005033
                     
      N. americanus G. inermis and H. lenta
Analysis of Differences   Property Mean Median SD Range Mean Median SD Range
    Depth 3.43 3 1.52 6.9-1.9 7.58 6.6 2.96 16.6-2.1
    Width 0.106 0.105 0.0157 0.14-0.082 2.48 2.3 0.515 4.3-1.7
    Height 0.0961 0.0923 0.0165 0.13-.078 2.33 2.3 0.391 3.2-1.6
    Circum 3.3 3.11 0.486 4.4-2.6 7.86 7.4 2.31 20.8-5.3
    Slope 62.4 70 22.6 90-25 79.6 85 11.3 90-45
    Compexity 2.4 2 0.737 4-2 2.21 2 0.463 4-2
    Tortuosity 6.56 1.23 14.0 43.0-1.0 1.02 1.0 0.124 1.4-0.5

TABLE 7. Matrix of Bray-Curtis test results for the comparison of Gorgyrella inermis and Hogna lenta burrows to G. inermis burrows. Cells are filled in black where burrows are compared to themselves. Cells with scores of high to moderate similarity (1.0-0.7) are filled in blue with increasing lightness. See Table 1 for burrow architecture key. (PDF only)

TABLE 8. Matrix of Bray-Curtis test results for the comparison of Gorgyrella inermis and Hogna lenta burrow to H. lenta burrows. Cells are filled in black where burrows are compared to themselves. Cells with scores of high to moderate similarity (1.0-0.7) are filled in blue with increasing lightness. See Table 1 for burrow architecture key. (PDF only)

TABLE 9. Mann-Whitney and Kolmogorov-Smirnov test results ( p values) from the comparison of clusters in each dendrogram. 1) Gorgyrella and Hogna (Figure 18), 2) Spiders and Aphonopelma (Figure 20), 3) Spiders and Hadrurus (Figure 21), 4) Spiders and Mastigoproctus (Figure 22), 5) Spiders and Narceus (Figure 23), 6) All arthropods (Figure 24). Boxed values indicate statistical difference ( p <0.05). All measurements are in cm, and all angles are in degrees.

1. Gorgyrella inermis and Hogna lenta
Clusters Test Depth Length Width Height W/H Ratio Circum Slope Complexity Tortuosity
A and B MW 0.5284 0.8672 0.6977 0.5268 0.2726 0.7476 1.83E-07 0.2561 0.136
KS 0.4373 0.8424 0.5739 0.4144 0.1505 0.4729 1.15E-08 0.005935 0.4033
                     
C and (A+B) MW 0.6862 0.7465 0.05439 0.03994 0.9541 0.04611 2.66E-02 0.1712 0.08435
KS 0.895 0.7386 0.03906 0.02582 0.7959 0.03184 2.58E-02 0.5583 0.02083
                     
Analysis of Differences: Property Mean Median SD Range Mean Median SD Range
A and B     Cluster A Cluster B
    Slope 86.4 87 3.92 90-75 68.9 70 5.84 75-60
    Complexity 2.23 2 0.425 3-2 2.08 2 0.277 3-2
C and (A+B)   Cluster C Cluster (A+B)
    Width 1.9 1.9 0 1.9-1.9 2.48 2.3 0.515 4.3-1.7
    Height 1.75 1.75 0.0707 1.9-1.9 2.37 2.35 0.517 3.2-1.6
    Circum 5.85 5.85 0.212 6.0-5.7 7.6 7.5 2.35 20.8-5.3
    Slope 59.75 59.75 6.72 64.5-55 81.3 85 9.26 90-60
    Tortuosity 0.8 0.8 0.283 1.0-0.6 1.02 1 0.124 1.4-05
                     
2. Spiders and Aphonopelma chalcodes
Clusters Test Depth Length Width Height W/H Ratio Circum Slope Complexity Tortuosity
A and B MW 0.8119 0.874 0.6901 0.6409 0.47 0.8031 5.25E-05 0.3396 0.3312
KS 0.9316 0.9316 0.7889 0.6404 0.1809 0.7559 2.30E-06 0.001312 0.2426
                     
C and (A+B) MW 0.7523 0.6274 0.04811 0.03615 0.8564 0.04138 3.67E-02 0.1424 0.07881
KS 0.8661 0.6369 0.0347 0.0238 0.8441 0.0288 4.16E-02 0.5559 0.01959
                     
D and (A+B+C) MW 0.0208 0.1911 0.2138 0.07124 0.941 0.3748 1.71E-02 0.5426 0.7274
KS 0.01583 0.2749 0.3897 0.04716 0.5551 0.737 1.30E-02 0.01918 0.6591
                     
Analysis of Differences: Property Mean Median SD Range Mean Median SD Range
A and B     Cluster A Cluster B
    Slope 86.4 87 3.92 90-75 68.9 70 5.84 75-60
    Complexity 2.226 2 0.425 3-2 2.08 2 0.277 3-2
C and (A+B)     Cluster C Cluster (A+B)
    Width 1.90 1.9 0 1.9-1.9 2.50 2.35 0.518 4.3-1.7
    Height 1.75 1.75 0.0707 1.8-1.7 2.37 2.32 0.377 3.2-1.6
    Circum 5.85 5.85 0.212 6-5.7 7.96 7.5 2.35 20.8-5.3
    Slope 59.8 59.8 6.71 64.5-55 81.3 85 9.260 90-60
    Tortuosity 0.8 0.8 0.283 2.0-1.6 1.02 1 0.124 1.4-0.05
D and (A+B+C)     Cluster D Cluster (A+B+C)
    Depth 2.6 2.6 0.707 3.1-2.1 7.76 6.65 3.65 16.6-2.5
    Height 1.75 1.75 0.0707 1.8-1.7 2.34 2.3 0.390 3.2-1.6
    Slope 35 35 14.1 45-25 80.3 85 10.1 90-55
    Complexity 2 2 0 2-2 2.22 2 0.767 4-2
                     
3. Spiders and Hadrurus arizonensis
Clusters Test Depth Length Width Height W/H Ratio Circum Slope Complexity Tortuosity
A and B MW 0.5284 0.8672 0.6977 0.5268 0.2726 0.7476 1.83E-07 0.2561 0.136
KS 0.4373 0.8424 0.5739 0.4144 0.1505 0.4729 1.15E-08 0.005935 0.4033
                     
C and (A+B) MW 0.6862 0.7465 0.05439 0.03994 0.9541 0.04611 2.66E-02 0.1712 0.08435
KS 0.895 0.7386 0.03906 0.02582 0.7959 0.03184 2.58E-02 0.5583 0.02083
                     
D and (A+B+C+G6) MW 0.0779 4.14E-07 3.54E-09 1.15E-04 5.29E-10 1.48E-08 1.10E-09 5.46E-09 8.05E-09
KS 0.09865 1.03E-05 2.70E-07 0.0009562 3.23E-12 7.68E-10 1.04E-11 3.92E-08 1.35E-06
                     
Analysis of Differences: Property Mean Median SD Range Mean Median SD Range
A and B     Cluster A Cluster B
    Slope 86.4 87 3.92 90-75 68.9 70 5.84 75-60
    Complexity 2.26 2 0.425 3-2 2.08 2 0.277 3-2
C and (A+B)     Cluster C Cluster (A+B)
    Width 1.9 1.9 0 1.9-1.9 2.5 2.35 0.377 3.2-1.6
    Height 1.75 1.75 0.0707 1.9-1.9 2.37 2.3 0.517 3.2-1.6
    Circum 5.85 5.85 0.212 6.0-5.7 7.6 7.5 2.35 20.8-5.3
    Slope 59.75 59.75 6.72 64.5-55 81.3 85 9.26 90-60
    Tortuosity 0.8 0.8 0.283 1.0-0.6 1.03 1.0 0.110 1.4-0.5
                     
D and (A+B+C+G6)   Cluster D Cluster (A+B+C+G6)
    Length 26.9 23 18.5 76-7.0 8.03 6.9 3.83 18.4-2.5
    Width 4.34 4.4 0.553 5.2-3.1 2.48 2.3 0.515 4.3-1.7
    Height 1.86 1.9 0.347 2.4-1.0 2.33 2.3 0.391 3.2-1.6
    W/H Ratio 2.41 2.4 0.434 3.1-1.3 1.06 1.0 0.117 1.5-0.9
    Circum 12.3 12.3 1.957 17.3-9.0 7.86 7.4 2.31 20.8-5.3
    Slope 17.2 15 11.7 45-5 79.6 85 11.3 90-45
    Complexity 4 3 2.12 9-3 2.21 2 0.463 4-2
    Tortuosity 1.96 1.3 1.94 9.0-1.1 1.02 1.0 0.124 1.4-0.5
                     
4. Spiders and Mastigoproctus giganteus
Clusters Test Depth Length Width Height W/H Ratio Circum Slope Complexity Tortuosity
A and B MW 0.5695 0.8012 0.883 0.6538 0.311 0.9417 4.21E-05 0.4108 0.3301
KS 0.6414 0.973 0.8003 0.8466 0.07347 0.9289 8.11E-06 4.49E-05 0.05098
                     
C and (A+B) MW 0.4613 0.07504 0.9264 0.09654 0.1914 0.9877 5.76E-03 0.1023 0.5669
KS 0.4241 0.2973 0.4528 0.1178 0.5131 0.4528 3.19E-03 0.2973 0.3445
                     
D and (A+B+C) MW 0.2556 7.86E-01 1.84E-01 5.40E-01 6.17E-02 1.69E-01 5.83E-03 5.12E-01 4.53E-01
KS 0.2853 8.30E-01 3.65E-01 0.4104 6.31E-02 3.65E-01 3.67E-03 4.63E-03 7.81E-01
                     
E and (A+B+C+D) MW 0.5828 4.62E-05 4.25E-04 3.03E-01 5.02E-05 6.58E-04 5.70E-06 6.66E-06 1.57E-06
KS 0.8059 7.21E-05 3.97E-04 0.2461 1.71E-05 8.79E-04 4.79E-07 8.79E-04 1.40E-06
                     
Analysis of Differences: Property Mean Median SD Range Mean Median SD Range
A and B     Cluster A Cluster B
    Slope 81.1 85 9.30 90-60 68.3 70 60.9 75-60
    Complexity 2.16 2 0.370 3-2 2.07 2 0.267 3-2
    Tortuosity 1.03 1.0 0.107 1.4-0.5 1.04 1.0 0.0514 1.1-1.0
C and (A+B)   Cluster C Cluster (A+B)
    Height 61.1 62.3 4.66 65-55 79.6 84.5 10.4 90-55
D and (A+B+C)     Cluster D Cluster (A+B+C)
    Length 6.73 8 2.64 8.5-3.7 8.39 7.4 4.58 26.5-2.5
    Width 2.87 3 0.321 3.1-2.5 2.62 2.5 0.707 5.8-1.7
    Height 2.13 1.9 0.404 2.6-1.9 2.26 2.3 0.447 3.2-1.0
    W/H Ratio 1.37 1.3 0.208 106-1.2 1.18 1.1 0.312 2.2-0.9
    Circum 9.73 10 2.21 11.8-7. 8.26 7.5 2.52 20.8-5.3
    Slope 48.3 50 2.89 50-45 77.9 83 12.4 90-45
    Complexity 2 2 0 2-2 2.19 2 0.441 4-2
    Tortuosity 1.05 1.06 0.0502 1.1-1.0 1.03 1 0.123 1.4-0.5
E and (A+B+C+D)   Cluster E Cluster (A+B+C+D)
    Length 22.2 19.25 18.0 66-12.5 8.40 7.4 4.58 26.5-2.5
    Width 4.16 4.15 1 5.7-2.4 2.62 2.5 0.707 5.8-1.7
    Height 2.13 1.95 0.12 2.9-1.6 2.26 2.3 0.447 3.2-1.0
    W/H Ratio 1.94 1.95 0.267 2.4-1.5 1.18 1.1 0.312 2.2-0.9
    Circum 12.2 10.9 3.09 17.0-8.3 8.26 7.5 2.52 20.8-5.3
    Slope 36.9 37.5 9.22 47-20 77.9 83 12.4 90-45
    Complexity 4.5 3.5 2.67 10-2 2.19 2 0.441 4-2
    Tortuosity 1.6 1.5 0.363 2.4-13 1.03 1.0 0.124 1.4-0.5
                     
5. Spiders and Narceus americanus
Clusters Test Depth Length Width Height W/H Ratio Circum Slope Complexity Tortuosity
A and B MW 0.775 0.9409 0.8151 0.6481 0.2242 0.8738 2.08E-08 0.2501 0.1065
KS 0.3038 0.3038 0.7461 0.5839 0.2052 0.6381 7.20E-10 2.85E-03 0.3431
                     
C and (A+B) MW 0.5194 0.7809 0.0084 0.006638 0.8771 0.0132 1.83E-03 0.5232 0.5072
KS 0.8792 0.726 0.009266 0.005264 0.6869 0.007009 1.13E-03 0.02982 0.3073
                     
D and (A+B+C+G6) MW 0.006464 6.31E-01 2.23E-02 4.36E-03 1.90E-02 8.97E-03 9.25E-05 4.53E-02 3.03E-03
KS 0.005894 2.10E-01 5.89E-03 0.002205 5.89E-03 5.89E-03 3.11E-05 1.72E-01 2.56E-02
                     
Analysis of Differences: Property Mean Median SD Range Mean Median SD Range
A and B     Cluster A Cluster B
    Slope 86.3 87 4.09 90-75 69.1 70 5.45 75-60
    Complexity 2.2 2 0.406 1.4-0.5 2.07 2 0.258 3-2
C and (A+B)     Cluster C Cluster (A+B)
    Width 0.955 0.955 1.09 1.9-0.0096 2.22 2.3 0.925 4.3-0.815
    Height 0.880 0.855 1.01 1.8-0.0083 2.10 2.3 0.825 3.2-0.0825
    Circum 4.47 4.41 1.59 6.1-3.1 7.40 7.35 2.69 20.8-2.6
    Slope 55.7 59.8 12.4 65-38.3 81.1 85 9.11 90-60
    Complexity 2.5 2 1.00 4-2 2.16 2 0.370 3-2
D and (A+B+C)     Cluster D Cluster (A+B+C)
    Depth 3.67 2.95 1.82 6.9-1.9 7.12 6.55 3.74 16.6-1.9
    Length 8.09 9.7 4.09 12.5-2.34 7.54 6.75 3.88 18.4-1.9
    Width 1.01 1.09 0.883 2.5-0.105 2.12 2.3 0.985 4.3-0.0096
    Height 0.841 0.878 0.695 1.9-0.0824 2.01 2.25 0.888 3.2-0.0083
    W/H Ratio 1.21 1.25 0.132 1.31-0.964 1.06 1.0 0.117 1.5-0.89
    Circum 4.04 3.4 1.67 7.4-3.1 7.18 7.3 2.73 20.8-2.6
    Slope 39.3 36.3 12.1 60-25 79.2 84.5 11.4 90-38.3
    Complexity 2.67 2.5 0.816 4-2 2.19 2 0.436 4-2
    Tortuosity 14.5 1.46 20.6 43-1.0 1.04 1.0 0.212 2.3-0.5
                     
6. All Arthropods
Clusters Test Depth Length Width Height W/H Ratio Circum Slope Complexity Tortuosity
A and B MW 0.6142 0.811 0.6131 0.9046 0.1495 0.6567 2.10E-09 0.6933 0.1103
KS 0.3741 0.212 0.6751 0.8472 0.1127 0.8173 3.00E-11 0.001851 0.2362
                     
C and (A+B) MW 0.6934 0.05814 0.8784 0.1142 0.4974 0.8288 8.54E-04 0.1377 0.1976
KS 0.559 0.1132 0.4343 0.1048 0.5988 0.3781 2.56E-04 0.07935 0.1176
                     
D and (A+B+C) MW 0.0873 0.5791 0.1606 0.008276 0.03095 0.164 5.36E-06 0.6551 0.005936
KS 0.1617 0.1538 0.02949 0.01397 0.004231 0.03811 1.00E-04 0.002128 0.1008
                     
E and F MW 0.004284 0.05873 0.6612 0.3658 0.022 0.8178 8.22E-06 0.0001995 0.3299
KS 0.002861 0.1523 0.3668 0.7977 0.08807 0.952 2.32E-06 0.0003 0.4693
                     
(A+B+C+D) and (E+F) MW 0.9072 3.51E-10 7.40E-11 0.2881 9.28E-10 4.38E-10 4.77E-14 3.44E-12 1.79E-07
KS 0.972 7.20E-09 2.25E-12 0.06005 2.41E-12 1.50E-11 1.74E-16 9.39E-11 1.07E-06
                     
Analysis of Differences: Property Mean Median SD Range Mean Median SD Range
A and B     Cluster A Cluster B
    Slope 85.9 85 4.09 90-75 68.9 70 5.77 75-60
    Complexity 2.17 2 0.381 3-2 2.17 2 0.529 4-2
C and (A+B)     Cluster C Cluster (A+B)
    Slope 61.9 64.5 4.39 65-55 80.9 85 9.05 90-60
D and (A+B+C)     Cluster D Cluster (A+B+C)
    Width 1.75 1.13 0.943 3.1-1.02 2.25 2.3 1.11 5.8-0.082
    Height 1.37 1.07 0.678 2.6-0.79 1.95 2.2 0.870 3.2-0.078
    W/H Ratio 1.26 1.28 0.177 1.6-0.965 1.16 1.10 1.10 2.2-0.89
    Circum 5.72 3.62 3.54 11.8-3.1 7.48 7.30 2.91 20.8-2.6
    Slope 43.6 41.7 9.29 60-33 79.4 83 10.2 90-55
    Complexity 2.25 2 0.469 3-2 2.21 2 0.481 4-2
    Tortuosity 11.1 1.16 18.5 43-1.0 1.06 1.0 0.209 2.3-0.5
E and F     Cluster E Cluster F
    Depth 5.22 4.2 3.25 16-2.8 7.85 7.7 2.57 12.4-2.9
    W/H Ratio 2.33 2.4 0.650 3.1-0.55 1.84 2 0.476 2.5-0.77
    Slope 13.3 13 6.58 25-5 35.9 35 7.84 47-25
    Complexity 2.87 3 0.352 3-2 5.38 4 2.63 10-3
(A+B+C+D) and (E+F)   Cluster (A+B+C+D) Cluster (E+F)
    Length 7.90 6.8 4.51 26.5-1.9 25.3 19 17.7 76-6.1
    Width 2.19 2.3 1.09 5.8-0.082 4.18 4.35 1.17 6.6-0.12
    W/H Ratio 1.17 1.1 0.287 2.2-0.89 2.10 2.05 0.617 3.1-0.55
    Circum 7.28 7.3 3.01 20.8-2.6 12.1 12.05 3.10 19.1-3.4
    Slope 75.4 80 15.2 90-33 23.8 22.5 13.5 47-5
    Complexity 2.21 2 0.476 4-2 4.04 3 2.19 10-2
    Tortuosity 2.10 1.0 6.67 43-0.5 1.75 1.3 1.55 9.0-0.52

TABLE 10. Mann-Whitney and Kolmogorov-Smirnov test results (p values) from the comparison of the properties of Gorgyrella and Hogna burrows produced under different experimental conditions including terrarium size, sediment density, and sediment moisture. No results indicated significant differences (p <0.05).

 

  Size: Sm to Lg Terrarium Density: Mod to Denser Density: Mod to Densest Moisture: Mod to Dry Moisture: Mod to Wet
G. inermis MW KS MW KS MW KS MW KS MW

KS

Depth 0.06619 0.05292 0.8852 0.5344 0.3123 0.5344 0.1939 0.1075 0.4875

0.7361

Total Length 0.1761 0.3572 0.665 0.5344 0.3123 0.5344 0.1939 0.1075 0.4875

0.7361

Averge Width 0.7133 0.2587 0.8852 0.9969 0.3123 0.2195 0.3123 0.1075 0.8170

0.9355

Average Height 0.9025 0.8757 0.6650 0.9969 0.3123 0.5344 0.3123 0.5344 0.8170

0.9356

Average Circ 0.7133 0.8778 0.1939 0.1075 0.3094 0.1075 0.4075 0.5344 0.8170

0.7361

Average Slope 0.8510 0.05292 0.1479 0.9969 0.5385 0.5344 0.7715 0.5344 1

0.7361

Complexity 0.1282 0.4772 0.1814 0.5344 0.8687 1 0.1814 0.5344 0.4017

0.7361

Tortuosity 0.2330 0.2235 0.6650 0.5344 0.1913 0.5344 0.1124 0.1075 0.1052 0.1376

 

 

 

 

 

 

 

 

 

 

 

  Size: Sm to Lg Terrarium Density: Mod to Denser Density: Mod to Densest Moisture: Mod to Dry Moisture: Mod to Wet
H. lenta MW KS MW KS MW KS MW KS MW KS
Depth 0.6625 0.9762 0.6625 0.3197 0.8597 0.9610 0.7728 0.9906 0.77828 0.4249
Total Length 0.3827 0.3197 0.6579 0.3197 0.8597 0.9610 0.7728 0.9906 0.3865 0.4249
Averge Width 0.6625 0.9762 0.6625 0.9762 0.8597 0.8214 0.7728 0.8979 0.3865 0.6001
Average Height 0.3827 0.3955 1 0.9762 0.8597 0.9610 0.7728 0.9906 0.7728 0.7796
Average Circ 1 0.9762 0.07652 0.03626 0.5926 0.6160 0.7671 0.7796 0.3743 0.4249
Average Slope 0.1642 0.3197 0.1642 0.3197 0.2756 0.1520 0.3173 0.4249 0.3329 0.4249
Complexity 1 0.9762 0.5050 0.9762 0.3865 0.9610 0.6831 0.9906 0.6831 0.9906
Tortuosity 0.6428 0.9762 0.3537 0.3197 0.8454 0.9988 0.6831 0.9906 0.7469 0.7796

 

TABLE 11. Mann-Whitney and Kolmogorov-Smirnov test results (p values) between burrows produced by Hogna lenta individuals with a brood and those without a brood. No results indicated significant differences ( p <0.05).

Dwelling vs. brood burrows of H. lenta  
Architecture Test Depth Total Length Avg Width Avg Height W/H Ratio Avg Circ Avg Slope Complexity Tortuosity
Vertical Shaft MW 0.0519 0.05019 0.8961 0.3608 0.8961 0.8961 0.3865 1 0.8724
KS 0.02626 0.02626 0.9208 0.1747 0.6509 0.9208 0.3658 0.3197 0.6509
                     
Subvertical Shaft MW 0.3286 0.3329 0.8465 0.8465 0.8465 0.8465 0.8407 1 0.8465
KS 0.4739 0.4739 0.7052 0.7052 0.9093 0.7052 0.9093 0.3197 0.7052