APPENDIX

List of valid fossil species of Opiliones, from oldest to most recent (primary source: Dunlop et al., 2018). (1)= Tetrophthalmi; (2)= Eupnoi; (3)= Dyspnoi; (4)= Laniatores; (5)= Cyphophthalmi; Bar= Bartonian; Bas= Bashkirian; Bur= Burdigalian; C= Carboniferous; Cal= Callovian; Cen= Cenomanian; D= Devonian; Ems= Emsian; Gzh= Gzhelian; J= Jurassic; K= Cretaceous; Lut= Lutetian; Mos= Moscovian; Ne= Neogene; Pa= Paleogene; Pra= Pragian; Vis= Visean; Ypr= Ypresian; *= Fossil genus.

Species Family/(Suborder) System Stage Outcrop/Region Type of fossilization
Eophalangium sheari Dunlop et al., 2004* Not described (1) D Pra-Ems Rhynie Chert
Brigantibunum listoni Dunlop and Anderson, 2005* Plesion (2) C Vis East Kirkton Limestones
Echinopustulatus samuelnelsoni Dunlop, 2004* Family indet. (3) C Mos Missouri Clay
Archaeometa nephilina Pocock, 1911 Archaeometidae (2) or (4) C Mos Measures of Coseley Clay
Arachnometa tuberculata Petrunkevitch, 1949 Family indet, (?) C Mos Measures of Coseley Clay
Hastocularis argus Garwood et al., 2014 Not described (1) C Bas Montceau-les-Mines Clay
Eotrogulus fayoli Thevenin, 1901* Eotrogulidae (3) C Bas Commentry Finely grained sandstones
Nemastomoides elaveris Thevenin, 1901* Nemastomoidae (3) C Bas Commentry Finely grained sandstones
Ameticos scolos Garwood et al., 2011 Family indet. (3) C Bas Montceau-les-Mines Clay
Macrogyion cronus Garwood et al., 2011 Plesion (2) C Bas Montceau-les-Mines Clay
Kustarachne tenuipes Scudder, 1890* Plesion (2) C Gzh Mazon Creek Clay
Nemastomoides longipes (Petrunkevitch, 1913) Nemastomoididae (3) C Gzh Mazon Creek Clay
Mesobunus martensi Huang, Selden and Dunlop, 2009* Sclerosomatidae (2) J Cal Daohugou Sandstones and lutites
Daohugopilio sheari Huang, Selden and Dunlop, 2009* Family indet. (2) J Cal Daohugou Sandstones and lutites
Mesobunus dunlopi Giribet et al., 2012 Sclerosomatididae (2) J Cal Daohugou Sandstones and lutites
Palaeosiro burmanicum Poinar, 2008 Stylocellidae (5) K Cen Myanmar Amber
Halitherses grimaldii Giribet and Dunlop, 2005* Halithersidae (3) K Cen Myanmar Amber
Petrobunoides sharmai Selden et al., 2016 Epedanidae (4) K Cen Myanmar Amber
Trogulus longipes Haupt, 1956 Trogulidae (3) Pa Lut-Ypr Geiseltal Lignite
Siro platypedibus Dunlop and Giribet, 2003 Sironidae (5) Pa Lut Bitterfeld Amber
Siro balticus Dunlop and Mitov, 2011 Sironidae (5) Pa Lut Baltic Region Amber
Caddo dentipalpus (Koch and Berendt, 1854) Caddidae (2) Pa Lut Baltic Region Amber
Dicranopalpus ramiger (Koch and Berendt, 1854) Phalangiidae (2) Pa Lut Baltic Region Amber
Opilio ovalis Koch and Berendt, 1854 ?Phalangiidae (2) Pa Lut Baltic Region Amber
Leiobunum longipes Menge, 1854 Sclerosomatidae (2) Pa Lut Baltic Region Amber
Sabacon claviger (Menge, 1854) Sabaconidae (3) Pa Lut Baltic Region Amber
?Histricostoma tuberculatum (Koch and Berendt, 1854) Nemastomatidae (3) Pa Lut Baltic Region Amber
?Mitostoma denticulatum (Koch and Berendt, 1854) Nemastomatidae (3) Pa Lut Baltic Region Amber
?Nemastoma incertum Koch and Berendt, 1854 Nemastomatidae (3) Pa Lut Baltic Region Amber
Proholoscotolemon nemastomoides (Koch and Berendt, 1854)* Cladonychidae (4) Pa Lut Baltic Region Amber
Piankhi steineri Dunlop, Bartel and Mitov, 2012* Family indet. (3) Pa Lut Baltic Region Amber
Lacinius bizleyi Mitov, Dunlop and Penney, 2015 Phalangiidae (2) Pa Lut Baltic Region Amber
Amilenus deltshevi Dunlop and Mitov, 2009 Phalangiidae (2) Pa Lut Bitterfeld Amber
Mitostoma gruberi Dunlop and Mitov, 2009 Nemastomatidae (3) Pa Lut Bitterfeld Amber
Stephanobunus mitovi Dunlop and Mammitzsh 2010 Phalangiidae (2) Pa Lut Baltic Region Amber
Petrunkevitchiana oculata (Petrunkevitch, 1922)* Family indet. (2) Pa Bar Florissant Lacustrine lutites
Amauropilio atavus (Cockerell, 1907) Sclerosomatidae (2) Pa Bar Florissant Lacustrine lutites
Amauropilio lacoei (Petrunkevitch, 1922) Sclerosomatidae (2) Pa Bar Florissant Lacustrine lutites
Philacarus hispaniolensis Cokendolpher and Poinar, 1992 Family indet. (4) Ne Bur Dominican Rep. Amber
Hummelinckiolus silhavyi Cokendolpher and Poinar, 1998 Samoidae (4) Ne Bur Dominican Rep. Amber
Pellobunus proavus Cokendolpher, 1987 Samoidae (4) Ne Bur Dominican Rep. Amber
Cosmobunus sagani nov. sp. (this study) Sclerosomatidae (2) Ne Bur Rubielos de Mora Lacustrine lutites