TABLE 1. Fossils of Cylindrotominae.

Name Sex Current designation Age (Ma) Locality/country Reference
Cylindrotoma biamoensis ? Valid 38.0-33.9 Biamo/Russia Freiwald and Krzemiński (1991)
Cylindrotoma borealis ? Valid 56.0-47.8 Fur/Denmark Freiwald (1991)
Cylindrotoma larssoni female? Valid 56.0-47.8 Fur Formation/Denmark Freiwald (1991)
Cylindrotoma monikae ? Valid 56.0-47.8 Fur Formation /Denmark Freiwald (1991)
Cyttaromyia veterana ? Not tipuloid 50.3-46.2 Green River/USA Cockerell (1921a); Brown (1988)
Cyttaromyia brevicornis   Limoniidae 38.0-33.9 Baltic amber Loew (1850); Krzemiński (1991a)
Cyttaromyia cancellata ? Cyttaromyia princetoniana 37.2-33.9 Florissant/USA Scudder (1894); Krzemiński (1991a)
Cyttaromyia clathrata ? Cyttaromyia princetoniana 37.2-33.9 Florissant/USA Scudder (1894); Krzemiński (1991a)
Cyttaromyia fenestrata ? Valid 37.2-33.9 Green River/USA Scudder (1877); Cockerell (1921a)
Cyttaromyia frelloi male Valid 38.0-33.9 Baltic amber Krzemiński (1998)
Cyttaromyia longicornis   Limoniidae 38.0-33.9 Baltic amber Loew (1850); Krzemiński (1991a)
Cyttaromyia longipes   Limoniidae 38.0-33.9 Baltic amber Loew (1850); Krzemiński (1991a)
Cyttaromyia lynnae female Valid 46.6-45.8 Kishenehn/USA This paper
Cyttaromyia obdurescens female Valid 50.3-46.2 Green River/USA Cockerell (1925)
Cyttaromyia oligocena ? Cyttaromyia princetoniana 37.2-33.9 Florissant/USA Scudder (1894); Krzemiński (1991a)
Cyttaromyia princetoniana ? Valid 37.2-33.9 Florissant/USA Scudder (1894)
Cyttaromyia quievreuxi male Valid 33.9-28.1 Middle Salt/France Séguy (1934)
Cyttaromyia rayona ? Valid 38.0-33.9 Biamo/Russia Freiwald and Krzemiński (1991)
Cyttaromyia reclusa male Valid 50.3-46.2 Green River/USA Cockerell (1925)
Cyttaromyia scudderi ? Valid 56.0-47.8 Ølst Formation/Denmark Freiwald (1991)
Cyttaromyia succini   Limoniidae 38.0-33.9 Baltic amber Loew (1850); Krzemiński (1991a)
Cyttaromyia vahldieki female Valid 56.0-47.8 Ølst Formation/Denmark Freiwald (1991)
Diogma gelhausi male Valid 38.0-33.9 Baltic amber Podenas (2000)
Oryctogma sackenii ? Limoniidae 37.2-33.9 Florissant/USA Scudder (1894); Evenhuis (1994)

 

 

TABLE 2. Families and species of the order Diptera from the 46 Ma Coal Creek Member of the Kishenehn Formation and the oldest described fossil of each family.

FamilyGenus speciesMuseum/NumberReferenceOldest known age
(Ma)
Reference
Tipulidae Tipula fji USNM 625687 This study 206-222 Krzemiński (1992)
Limoniidae Helius constenius
Ellipteroides kishenehn
CMNH 34935
USNM 621123
Krzemiński (1991b)
This study
  Krzemiński (1992)
Cylindrotomidae Cyttaromyia lynnae USNM 621109 This study 49-56 Freiwald (1991)
Psychodidae Bruchomyiinae sp. USNM 619952 This study 182-183 Ansorge (1994)
Dixidae Dixella intacta
Dixella eomarginata
Dixella curvistyla
Dixella spinilobata
USNM 546204
USNM 553521
USNM 546860
USNM 553522
Greenwalt and Moulton (2016) 190-202 Lukashevich (1996)
Culicidae Culiseta kishenehn
Culiseta lemniscata
Culiseta? sp.
USNM 546528
USNM 547065
USNM 559050
Harbach and Greenwalt (2012); Greenwalt et al. (2013) 94-100 Borkent and Grimaldi (2011)
Anisopodidae Sylvicola silibrarius USNM 626077 This study 190-202 Rohdendorf (1962); Krzemińska et al. (2010)
Scatopsidae Efcookella nigra USNM 618088 This study 94-100 Fate et al. (2013)
Bibionidae Bibiodes kishenehnensis USNM 625738 This study 123-126 Ren et al. (1995)
Sciaridae Eosciarites hermes USNM 624633 This study 100-113 Kaddumi (2007)
Mycetophilidae Rymosia hypnolithica USNM 624134 This study 122-125 Ren et al. (1995)
Bolitophilidae Bolitophila warreni USNM 595138 Greenwalt and Blagoderov (2019) 46* Greenwalt and Blagoderov (2019)
Rhagionidae Litoleptis araeostylus USNM 624657 This study 242-247 Krzemiński and Krzemińska (2003)
Bombyliidae Eoanomala melas USNM 595155 Greenwalt et al. (2015b) 56-59 Nel (2008)
Asilidae Kishenehnoasilus bhl USNM 624491 This study 113-123 Grimaldi (1990)
Dolichopodidae Salishomyia eocenica USNM 622501 This study 125-130 Grimaldi and Cumming (1999)
Hybotidae Drapetis adelomedos USNM 621705 This study 125-130 Hennig (1970)
Platypezidae Agathomyia eocenica USNM 553697 This study 113-126 Mostovski (1995)
Lonchopteridae Lonchoptera eocenica USNM 625379 This study 129 Grimaldi and Cumming (1999)
Phoridae Aenigmatias kishenehnensis USNM 625132 Thu study 46 This study
Pipunculidae Pipunculinae sp. USNM 625934 This study 48-56 Archibald et al. (2014a)

 

 

TABLE 3. Families of the order Diptera from four North American Compression fossil Lagerstätten.

Dipteran familyKishenehnG. RiverFlorissantOkanagan
Agromyzidae   x x  
Anisopodidae x x    
Asilidae x x x  
Bibionidae x x x x
Blephariceridae   x    
Bolitophilidae x     x
Bombyliidae x   x  
Cecidomyiidae   x x  
Chironomidae   x x  
Conopidae   x    
Culicidae x x    
Cylindrotomidae x x x  
Dixidae x      
Dolichopodidae x   x  
Empididae   x x x
Eophlebomyiidae   x    
Glossinidae     x  
Heleomyzidae   x x  
Hybotidae x      
Keroplatidae   x   x?
Lauxaniidae     x  
Limoniidae x x x  
Lonchopteridae x      
Lygistorrhyinidae       x?
Muscidae   x x  
Mycetophilidae x x x x
Mydidae     x  
Mythicomyiidae   x    
Nemestrinidae     x  
Oestridae   x    
Phoridae x   x  
Piophilidae     x  
Pipunculidae x   x x
Platypezidae x x x  
Psychodidae x     x
Ptychopteridae     x x
Rhagionidae x   x x
Richardiidae     x  
Scathophagidae     x  
Scatopsidae x   x  
Sciaridae x x x x
Sciomyzidae   x x x
Sepsidae     x  
Stratiomyidae   x x  
Syrphidae   x x x
Tabanidae     x  
Tachinidae   x    
Therevidae     x  
Tipulidae x x x x
Trichoceridae       x?
Ulidiidae     x  
Xylomyidae     x