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TABLE 1. Early Miocene sites from Spain with remains of shrews and heterosoricids, excluding this work (Agustí et al., 1988; Van der Meulen, 2003; Van den Hoek Ostende and Furió, 2005; Van den Hoek Ostende et al., 2009; 2017; Van der Meulen et al., 2012; Furió et al., 2018; Jovells-Vaquè et al., 2018).

Taxon Basin Site/s
Soricidae indet. Almazán Cetina de Aragón
  Calatayud-Montalbán Ateca 1, 3; Bañon 5; Moratilla 1; Olmo Redondo 3; Valtorres
  Ebro La Nasa 1, 6; Mediavilla; Rincón del Bu
  Madrid O’Donnell
  Rubielos de Mora Rubielos de Mora 1, 2
  Teruel Montalvos 2
Oligosorex (cf. or aff.) thauensis (Crochet, 1975) Calatayud-Montalbán Agreda; Bañon 2; Moratilla 1; Ramblar 1, 3, 4, 7; San Roque 4a
  Ebro Barranco del Fraile; Cabezo Vaquero; Fuenmayor 2; Valhondo 1
  Rubielos de Mora Alto de Ballester 1
Oligosorex sp. Calatayud-Montalbán Artesilla, Olmo Redondo 1, 2 and 3, San Marcos, San Roque 2 and 3, Villafeliche 2a, Vargas 1A, 2A, 4A, 4B and 4BB
  Magro Buñol
Clapasorex (cf.) alvarezae van den Hoek Ostende, 2003 Calatayud-Montalbán Navarrete del Río, Ramblar 1 and 3, Valhondo 1
  Rubielos de Mora Alto de Ballester 1
Soricella discrepans Doben-Florin, 1964 Calatayud-Montalbán Navarrete del Río, Ramblar 1
Miosorex grivensis (Deperet, 1892) Calatayud-Montalbán Munebrega 1, San Marcos, Torralba 1, Vargas 4A and 4B
Florinia ” Calatayud-Montalbán Olmo Redondo 2, San Roque 1, 2, 3, 4a and 5, Vargas 1A, 2A, 4A, 4B and 4BB
Paenelimnoecus (cf.) truyolsi (Gibert, 1975) Calatayud-Montalbán San Marcos, Vargas 1A
  Rubielos de Mora Alto de Ballester 1
Heterosorex sp. Almazán Cetina de Aragón
  Calatayud-Montalbán Artesilla, Navarrete del Río, Ramblar 1, Vargas 1A, 3, 4A and 4BB
  Ebro Alto de Ballester 1
  Madrid O’Donnell
Heterosorex (cf.) neumayrianus (Schlosser, 1887) Magro Buñol
  Ribesalbes-Alcora Mas de Antolino 2
  Teruel Montalvos 2
  Vallès-Penedès Les Cases de la Valenciana
Heterosorex ruemkeae Doukas, 1986 Magro Buñol

 

 

 

 

TABLE 2. Measurements of the teeth of soricids and heterosoricids from the Ribesalbes-Alcora Basin. AW: Anterior width; BL: Buccal length; H: Height; L: Length; LL: Lingual length; LT: Length of the talon; PE: Posterior emargination; PW: Posterior width; TAW: Talonid length; TRW: Trigonid length; W: Width.

Element Site Measurement n Min. Med. Max.
Oligosorex thauensis
i MAB5 L 1   2.10  
    W 2 0.58 0.63 0.68
  MAB11 L 1   1.81  
    W 1   0.79  
a2 MAB3 L 1   0.84  
    W 1   0.58  
p4 MAB11 L 1   1.06  
    W 1   0.69  
m1 MAB0B L 2 1.17 1.20 1.22
    TRW 2 0.76 0.78 0.80
    TALW 2 0.81 0.82 0.83
  MAB5 L 1   1.24  
    TRW 1   0.76  
    TALW 1   0.77  
  MAB11 L 2 1.21 1.24 1.27
    TRW 2 0.78 0,79 0.80
    TALW 3 0.77 0.79 0.81
  MAB11B L 1   1.23  
    TRW 1   0.61  
    TALW 1   0.74  
  CBR0D L 1   1.28  
    TRW 1   0.79  
    TALW 1   0.86  
m2 MTR2 L 1   1.16  
    TRW 1   0.71  
    TALW 1   0.76  
  MAB0B TALW 1   0.8  
  MAB3 L 1   1.17  
    TRW 1   0.72  
    TALW 1   0.79  
  MAB5 L 1   1.10  
    TRW 1   0.69  
    TALW 1   0.69  
  MAB11 L 1   1.28  
    TRW 1   0.73  
    TALW 1   0.84  
m1,2 MTR3 TALW 1   0.76  
  MAB11A TALW 1   0.81  
m3 MAB3 L 1   0.91  
    W 1   0.75  
  MAB5 L 4 0.96 1.01 1.05
    W 3 0.62 0.70 0.77
  MAB11 L 1   1.06  
    W 1   0.73  
  CBR0D L 1   1.04  
    W 1   0.64  
I MAB0B L 1   1.18  
    LT 1   0.52  
    H 1   0.96  
  MAB3 Length 1   1.13  
    LT 2 0.51 0.53 0.55
    H 3 0.85 0.89 0.93
  MAB5 Length 1   1.23  
    LT 1    0.60  
    H 1   0.85  
  MAB11 Length 1   1.23  
    LT 2 0.55 0.58 0.61
    H 2 0.91 0.93 0.94
A1 MAB3 Length 1   0.99  
    Width 1   0.73  
P4 MAB3 BL 1   1.44  
    Width 1   1.08  
    PE 1   0.90  
    LL 1   0.95  
  MAB5 BL 4 1.36 1.39 1.44
    Width 2 1.18 1.23 1.28
    PE 2 0.78 0.85 0.91
    LL 1   0.91  
  MAB11 BL 1   1.58  
M1 MAB3 BL 2 1.21 1.23 1.24
    PE 1   0.94  
    AW 2 1.16 1.22 1.27
  MAB5 BL 2 1.18 1.19 1.19
    PE 2 0.89 0.92 0.94
    LL 2 1.16 1.18 1.19
    AW 2 1.19 1.23 1.27
    PW 2 1.49 1.50 1.51
  MAB11 BL 1   1.27  
  MAB11B PE 1   0.89  
    LL 1   1.20  
    AW 1   1.29  
M2 MAB0A BL 1   1.20  
    PE 1   0.92  
    LL 1   1.12  
    AW 1   1.48  
    PW 1   1.48  
  MAB3 BL 2 1.15 1.19 1.22
    PE 3 0.89 0.91 0.93
    LL 2 1.08 1.15 1.22
    AW 1   1.43  
  MAB5 BL 1   1.07  
    PE 1   0.89  
    LL 1   1.22  
    AW 2 1.30 1.33 1.35
  CBR0G BL 1   1.13  
    PE 1   0.84  
    LL 1   1.06  
    AW 1   1.39  
    PW 1   1.02  
M3 MAB0B Length 1   0.68  
    Width 1   1.14  
  MAB3 Length 1   0.71  
    Width 1   1.23  
  MAB5 Length 4 0.77 0.83 0.92
    Width 3 0.96 1.03 1.10
cf. Soricella discrepans
M1 FS1 PE 1   1.02  
Paenelimnoecus micromorphus
i BC1 Width 1   0.54  
p4 BC1 Length 1   0.67  
    Width 1   0.55  
m1 BC1 L 2 1.01 1.05 1.09
    TRW 2 0.67 0.67 0.67
    TAW 2 0.72 0.73 0.73
m2 BC1 L 2 1.07 1.09 1.10
    TRW 2 0.69 0.70 0.70
    TAW 2 0.73 0.74 0.74
m3 BC1 L 2 0.89 0.90 0.91
    W 2 0.52 0.56 0.60
I BC1 LT 1   0.49  
    H 1   0.82  
M2 BC1 PE 1   0.79  
    LL 1   1.05  
Heterosorex neumayrianus
i MAB3 L 1   6.25  
    W 1   1.80  
a3 BC1 L 1   0.65  
    W 1   0.76  
p4 MAB3 L 1   1.03  
    W 1   0.98  
m1 MTR2 L 1   1.99  
    TRW 1   1.36  
    TAW 1   1.40  
  MAB3 L 1   1.87  
    TRW 2 1.26 1.27 1.27
    TAW 1   1.32  
m2 MAB3 L 1   1.60  
    TRW 1   1.37  
    TAW 1   1.29  
m1,2 MAB3 TAW 1   1.39  
I MAB3 L 1   2.83  
    LT 1   0.93  
    H 1   1.87  
A1 MAB0B L 2 1.65 1.67 1.68
    W 2 1.31 1.37 1.43
  MAB3 L 1   1.74  
    W 1   1.35  
  MAB5 L 2 1.66 1.74 1.81
    W 2 1.36 1.39 1.43
A2 MAB3 L 3 1.08 1.13 1.19
    W 3 0.90 0.93 0.97
A3 MTR2 L 1   1.00  
    W 1   0.81  
A4 MAB0B L 1   0.92  
    W 1   0.61  
P4 MAB5 BL 2 1.54 1.57 1.59
    W 2 1.53 1.54 1.55
    PE 1   1.24  
    LL 1   1.12  
M1 MAB3 BL 1   1.73  
    PE 1   1.80  
    LL 1   1.75  
    AW 1   1.80  
    PW 1   1.82  
M3 MAB11B L 1   0.82  
    W 1   1.33  

 

 

TABLE 3. Percentages of frequency (%) of soricids, heterosoricids, dimylids, talpids and erinaceids found in the Araia sections. Only represented are the sites intensively sampled with more than 50 teeth, the number in parentheses next to the locality name represents the number of mammal teeth found at this site, and the number in parenthesis next to the frequency is the percentatge of the family with respect to the mammals teeth found in the sites studied from Ribesalbes-Alcora Basin (modified from Crespo et al., 2019).

Locality Soricids Heterosoricids Dimylids Talpids Erinaceids
CBR0B (56) 0 16.8 (1.8) 0 83.2 (8.9) 0
MAB11 (145) 52 (11.9) 0 3.1 (0.7) 24 (5.5) 20.9 (4.8)
MAB5 (874) 12.3 (2.7) 4.1 (0.9) 51.4 (11.3) 15.5 (3.4) 16.8 (3.7)
MAB3 (807) 25.5 (2.4) 18.1 (1.7) 0 24.5 (2.3) 31.9 (3)
FS1 (63) 25 (1.6) 0 0 25 (1.6) 50 (3.2)
MAB0B (75) 33.3 (8) 16.7 (4) 0 38.7 (9.3) 11.3 (2.7)
MAB0A (80) 33.3 (1.3) 0 0 33.3 (1.3) 33.3 (1.3)
BC1 (180) 48.3 (7.2) 7.4 (1.1) 7.4 (1.1) 29.5 (4.4) 7.4 (1.1)
MTR2 (187) 5.8 (0.5) 18.6 (1.6) 0 50 (4.3) 25.6 (2.2)
MCX3 (132) 0 0 0 9.5 (0.8) 90.5 (7.6)

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