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TABLE 1. Taxonomic study history of palaeolenid trilobites from the Guanshan Biota.

Publication Species considered Relevant notes
Mansuy (1912, p. 27) Palaeolenus douvillei, P. deprati and P. “lantenoisi” First description of three Palaeolenus species using a few specimens (n=6) with all the cranidial features exhibiting
Chang (1966, p. 150). Megapalaeolenus deprati Moved Palaeolenus deprati into Megapalaeolenus
Chang et al. (1980, p. 234, pl. 73, Figs 9-12) Palaeolenus longispinus Erected Palaeolenus longispinus using robust pleural spines on second thoracic tergite
Lin and Peng (2004) Megapalaeolenus deprati and Palaeolenus douvillei Doubted the validity of Megapalaeolenus. Noted that holotype was deformed and considered M. deprati conspecific with Palaeolenus douvillei
Luo et al. (2007) Megapalaeolenus deprati, Palaeolenus douvillei and P. “lantenoisi” Considered all species and genera to be valid
Lin and Peng (2009) Megapalaeolenus deprati and Palaeolenus douvillei Considered species conspecific
Hu et al. (2013) Megapalaeolenus deprati, Palaeolenus douvillei, P. “lantenoisi” and P. longispinus Suggested all taxa were valid species
Lin (2014) All the published 34 specimens assigned to palaeolenids (n=34) Used cluster analysis to group all taxa into Palaeolenus. Suggested P. douvillei and Megapalaeolenus deprati conspecific
Luo et al. (2014) Palaeolenus douvillei and Megapalaeolenus deprati Considered both taxa valid. Noted they are found in different members of the Wulongqing Formation
Lin and Peng (2016) All the published 34 specimens assigned to palaeolenids (n=34) Used cluster analysis to group. Reconfirmed Lin (2014), suggesting Palaeolenus douvillei and Megapalaeolenus deprati are conspecific

 

 

TABLE 2. Locations of cephalic landmarks for analyzed specimens.

Landmark Description
1 Intersection of anterior border with sagittal line
2 Intersection of border furrow with sagittal line
3 Intersection of anterior margin of frontal lobe with sagittal line
4 Intersection of anterior margin of occipital ring with sagittal line
5 Intersection of posterior margin of occipital ring with sagittal line
6 Intersection of axial glabellar furrow and posterior margin
7 Anterior border of palpebral lobe
8 Posterior border of palpebral lobe

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