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TABLE 1. Measurements and proportions of specimens of Armigatus carrenoae sp. nov. from the Tlayúa quarry referred here. All the measurements are given in millimeters, and all percentages are calculated on the base of the standard length (SL).

  IGM 11536 IGM 11537 IGM 11538 Measurements
(in mm) and proportions
IGM 11539
IGM 11540 IGM 11541 IGM 1542 IGM 11543 Avg. (Range)
Total length 57.2 68.3 57 81.9? 35.3 57.5 56.6? 68.3? 55.6 (35.3-68.3)
Standard length 42.7 50.1 42.4 64.2? 49.7 46.0 46.9? 51.8 47.1 (18.4-51.8)
Body height
% SL
23.0
53.8
23.7
47.3
22.7
53.5
27.3
42.5?
30.6?
61.5?
21.7
47.1
25.3?
53.4?
25.3?
53.4?
24.4 (23-28.2)
51.2 (47.1-54.4)
Head length
% SL
17.2
40.2
18.2
37.5
16.7
39.3
22.5
35?
18.4
37.0
16.5
35.8
17.8?
37.9?
18.4?
35.5?
18.25 (18.4-22.5)
37.9 (35.8-40.2)
Predorsal length
% SL
24.2
56.6
28.5
56.8
24.4
57.5
34.9?
54.3?
28.2
56.7
25.8
57.8
27.6?
58.8?
28.4
54.8
26.5 (24.2-28.5)
56.7 (54.8-57.8)
Dorsal fin length
% SL
7.0
16.3
8.2
16.3
6.5
15.3
9.5?
16.2?
7.4
14.8
8.0
17.3
7.6
16.2?
8.3
17
7.4 (6.7-8.3)
16.1 (14.8-17.3)
Prepelvic length
% SL
25.2
59
30.3
60.4
25.2
59.4
35?
54.5?
29.6
60.2
26.6
57.8
26?
55.4?
29.4
56.8
27.7 (25.2-30.3)
58.9 (56.8-60.4)
Preanal length
% SL
33.7
78.9
38.5
76.8
33.4
78.7
52.6?
81.9?
40.0
80.4
35.3
76.7
37.5?
79.9?
42.2
81.4
37.1 (33.4-42.2)
78.8 (76.7-81.4)
Anal fin length
% SL
8.1
18.9
8.3
16.5
7.3
17.2
10.9
16.9?
8.5
17.1
8.4
18.2
7.8
15.3?
?
?
8.5 (7.3-10.9
17.5 (16.5-18.9)
          Meristics       Mode (Range)
Total vertebrae 35 33 34 34 35 35 33+ 29+ 34 (33-35)
Abdominal centra 17 16 15 16 16 16 14+ 10+ 16 (15-17)
Caudal centra 16 15 17 16 17 17 17 17 17 (15-17)
Pair of ribs 14 15 15 15 14 14 15 ? 14-15
Predorsal scutes 8 5+ 7? 3+ 3+ 4+ 5+ 3+ 8
Abdominal scutes 22? 23 22 18+ 22? 21+ 23 5+ 23 (22-23)
Prepelvic scutes 16? 17 16 12+ 16? 15+ 17 5+ 17 (16-17)
Postpelvic scutes 6 6 6 6 6 6 6 ? 6
Dorsal fin rays ii, 11 i, 11 ii, 11 ii,? ii, 9? ii,13 ii, 11 ii,11 ii,11 (i-ii,11-13)
Dorsal pts. 11 11 11 11 10+ 12 9+ ? 11 (11-12)
Anal fin rays ii, 17 ii, 15 ii, 17 ii, 17 I, 14? ii, 17 ?, 5+ ii, 17? ii, 17 (ii,15-17)
Anal pts. 17 15 16 15 14+ 17 14+ 12+ 17 (16-18)
Pectoral fin rays 15 15 14+ 13+ 14 5+ 14 15 15 (14-15)
Pelvic fin rays 6 5 5 5 5? 5 7 6+ 5(5-7)
Predorsal bones 6 6 6 6 6 6 6 5? 6

 

 

TABLE 2. Nominal species currently included in the order Ellimmichthyiformes.

Species Age and Distribution
Armigatus alticorpus Forey, Yi, Patterson, and Davies, 2003. Cenomanian. Namoura, Lebanon.
Armigatus brevissimus (Blainville, 1818). Cenomanian. Heckel and Hajula, Lebanon.
Armigatus carrenoae sp. nov. (present manuscript). Albian. Tlayúa quarry, Mexico.
Armigatus dalmaticus Murray, Vernygora, Japundžić, Radovčić, Wilson, Bardack, and Grande, 2016. Campanian? Dalmatia, Croatia.
Armigatus namourensis Forey, Yi, Patterson, and Davies, 2003. Cenomanian. Namoura, Lebanon.
Armigatus oligodentatus Vernygora and Murray, 2016. Cenomanian/Turonian. Agoult, Morocco.
Codoichthys carnavalii Silva Santos, 1994. Aptian. São Luís-Grajaú Basin, Brazil.
Diplomystus altisomus Yabumoto, 1994 Early Cretaceous. Kyushu, Japan.
Diplomystus birdi Woodward, 1895. Cenomanian. Heckel and Hajula, Lebanon.
Diplomystus dentatus Cope, 1877. Eocene. Green River Fm., USA.
Diplomystus dubertreti Signeux, 1951. Santonian. Sahel Alma, Lebanon.
Diplomystus kokuraensis Uyeno, 1979. Neocomian. Kyushu, Japan.
Diplomystus primotinus Uyeno, 1979. Neocomian. Kyushu, Japan.
Diplomystus solignaci Gaudant and Gaudant, 1971. Senonian. Gabès, Tunisia.
Diplomystys shengliensis Zhang, Zhou, and Qing, 1985. Eocene. Shandong, China.
“Diplomystus” trebicianensis Bannikov and Sorbini, 2000. Paleocene, Trebiano, Italy.
Ellimma branneri (Jordan, 1910). Hauterivian-Aptian. Muribeca Fm., Brazil.
Ellimma cruzi Silva Santos 1990. Aptian. Cabo Fm., Brazil.
Ellimma longipectoralis Polck, Gallo, Figueiredo, Viana, Santos, Quiroz-Neto, and Jahnert, 2020. Aptian. Velha Fm., Santos Basin, Brazil.
Ellimmichthys longicostatus (Cope, 1886). Hauterivian-Barremian. Recôncavo Fm., Brazil.
Ellimmichthys goodi Eastman, 1912. Aptian-Albian. Cocobeach Series, Equatorial Guinea.
Ellimmichthys maceioensis Malabarba, Do Carmo, Gómez-Pérez, and Queiroz-Neto, 2004. Aptian-Albian. Maceio Fm., Brazil.
Eoellimmichthys superstes Marramà, Bannikov, Kriwet, and Carnevale, 2019. Early Eocene. Pesciara, Italy.
Ezkutuberezi carmenae (Poyato-Ariza, López-Horgue, and García-Garmilla, 2000). Valanginian-Barremian. Villaro Fm., Spain.
Foreyclupea loonensis Vernygora, Murray and Wilson, 2016. Albian. Loon River Fm., Canada.
Gateroclupea branisai Signeux, 1964 (see Marramà and Carnevale, 2017). Late Cretaceous-Paleocene. Cayara, Bolivia.
Horseshoeichthys armaserratus Newbrey, Murray, Brinkman, Wilson, and Neuman, 2010. Maastrichtian. Horseshoe Canyon Fm., Canada.
Kwangoclupea dartevellei (Casier, 1965). Cenomanian. Kwango, Zaire.
Ornategulum sardinoides (Pictet, 1850). Cenomanian. Heckel and Hajula, Lebanon.
Paraclupea chetungensis Sun, 1956. Early Cretaceous. Chawan Fm., China.
Paraclupea seilacheri Alvarado-Ortega and Melgarejo-Damián, 2017. Albian. Tlayúa, Mexico.
Ranulfoichthys dorsonudum Alvarado-Ortega, 2014. Albian. Tlayúa, Mexico.
Rhombichthys intoccabilis Khalloufi, Zaragüeta-Bagils, and Lelièvre, 2010. Cenomanian. Ein Yabrud, Palestine.
Scutatoclupea bacchiai Bannikov, 2015. Cenomanian. Namoura, Lebanon.
Scutatuspinosus itapagipensis Santos and Corrêa, 1985. Hauterivian?. Marfim Fm., Brazil.
Sorbinichthys elusive Bannikov and Bacchia, 2000. Cenomanian. Namoura, Lebanon.
Sorbinichthys africanus Murray and Wilson, 2011. Cenomanian/Turonian. Agoult, Morocco.
Thorectichthys rhadinus Murray and Wilson, 2013. Cenomanian/Turonian. Agoult, Morocco.
Thorectichthys marocensis Murray and Wilson, 2013. Cenomanian/Turonian. Agoult, Morocco.
Triplomystus applegatei Alvarado-Ortega and Ovalles-Damián, 2008. Cenomanian. El Espinal, Mexico.
Triplomystus noorae Forey, Yi, Patterson, and Davies, 2003. Cenomanian. Namoura, Lebanon.
Triplomystus oligocostatus Forey, Yi, Patterson, and Davies, 2003. Cenomanian. Namoura, Lebanon.
Tycheroichthys dunveganensis Hay, Cumbaa, Murray, and Plint, 2007. Cenomanian. Dunvegan Fm., Canada.
Tunisiaclupea speratus Boukhalfa, Wu, Ben Ali, and Fang, 2019. Late Barremian. Chotts Basin, Tunisia.

 

TABLE 3. Comparison of the nominal species of the genus Armigatus. Maximum standard length (SL) expressed in millimeters, (based on data from table 3; Forey et al., 2003; Murray et al., 2016, tables 2, 3; Vernygora and Murray, 2016, table 1; Vernygora and Murray, 2021).

Species→
Number of individuals considered→
Armigatus carrenoae sp. nov.
(8)
Armigatus brevissimus
(10)
Armigatus namouraensis
(11)
Armigatus alticorpus
(6)
Armigatus oligodentatus
(8)
Armigatus dalmaticus
(6)
Averages of proportions
expressed as % of SL (ranges)
Maximum TL ≈ 81.9 105 139 47 37 48.6
Body height 51.2
(47.1-54.4)
37.3
(33-42)
37.7
(31-50)
48.9
(46.1-52.7)
32.1
(28-36)
40.8
(34.6-44-8)
Head length 37.9
(35.8-40.2)
33.6
(30-38)
31.9
(28-35)
34.2
(33.3-36.1)
32.5
(31-35)
31.3
(28-33)
Predorsal length 56.7
(54.8-57.8)
46.5
(44-49)
47.3
(43-52)
49.2
(47.9-51)
48.6
(46-52)
46.3
(43.3-50.2)
Preanal length 78.8
(76.7-81.4)
67.1
(64-72)
69.1
(66-77)
69.9
(66.6-73.7)
77.8
(75-82)
75.4
(66.5-81.9)
Prepelvic length 58.9
(56.8-60.4)
50
(47-56)
51.5
(49-56)
53.6
(55.5-50)
55.2
(50-58)
56.4
(55.4-57.8)
Modes of accounts
(ranges)
Anal fin rays ii, 17
(ii,15-17)
28
-
25
(24-27)
25+
(?-25+)
?
(12?-14?)
16
-
Dorsal fin rays ii,11
(i-ii,11-13)
19
-
19
(18-19)
16
(15-17)
11
(10?-13?)
13, 14
(13-14)
Pectoral fin rays 15
(14-15)
20
(18-20)
12
(12-15)
12 or 13?
(12-13)
8
(8-13)
?
(12-13)
Pelvic fin rays 5
(5-7)
7
-
8
(8-9)
?
-
7
(6-9)
7?
-
Predorsal bones 6
(6)
6
(6)
6
(5-6)
6
(5-6)
8
(7?-8)
7, 8
(7-8)
Total vertebrae

Abdominal

Caudal (including urals)
34
(33-35)
16
(15-17)
19
(17-19)
31
(31-32)
17
(16-18)
15
(13-15)
38
(36-38)
18
(17-19)
20
(17-21)
36 or 37
(35-37)
16
(16-18)
20
(19-21)
38
(37-39)
24?
(23-25)
14, 15
(14-15)
34
(32-37)
¿
(14-19)
18
(13?-18)
Predorsal scutes 8
(8)
9
(9)
17
(17-19)
11?
(7+-11?)
?
(5?-9?)
11
(10-11)
Prepelvic scutes 17
(16-17)
16
-
20
(19-21)
17
-
13
(10-13)
13
(13)
Postpelvic scutes
(6)
7
(6?-7)
8
(8-9)
7
(7)
10, 11
(8-11)
9-10
(9-10)
Other
features
Skull roof bones Stout ridges ornament the frontal, parietal, and pterotic. Smooth
Parasphenoid with the osteoglossid-like tooth patch ? toothless
Opercle ornaments Basal short and straight grooves ? Smooth
Epineurals and epipleurals close to caudal peduncle far from the caudal peduncle close to caudal peduncle far from the caudal peduncle
Predorsal bones below predorsal scutes Posterior 4-3 (= predorsal series inccomplete All (= predorsal series complete) Posterior 4 ? All (= predorsal series complete)
Posterior
predorsal scutes
Posterior ones subtriangular, others are ovoid, all with spine Posterior ones subtriangular with a small spine, others ahead become heart-shaped All are heart-shaped All are wide and rectangular

With no posterior spine

All are heart-shaped All subrectangular with anterior and posterior spines

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