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MATERIALS
Recent and fossil specimens examined for this study belong to: Geological Institute of the Siberian Branch RAS, Ulan-Ude; Institute of Plant and Animal Ecology USC, Yekaterinburg; Zoological Institute RAS, Saint Petersburg, Russia and Institute of Geology SB RAS, Yakutsk (Russia); American Museum of Natural History, New York; Smithsonian Institution, Washington, D.C.; Carnegie Museum of Natural History, Pittsburgh; Denver Museum of Nature and Science; and the University of California, Museum of Paleontology (USA).
The systematic organization of fossil species presented below is based on the morphology of preserved skulls, mandibles and teeth, and on biometric studies. Most living species, in addition to the mentioned characters, were diagnosed according to external features such as body ratio or pelage. The comparative analysis of living and extinct species is based mainly on the tooth structures and in part on preserved fragments of skull and mandibles for fossil forms.
We follow the standard Ochotona dental morphology outlined by
Erbajeva (1988). In a whole more than 2000 specimens of living pikas and approximately 650 specimens of fossil Ochotona were examined from mentioned above Institutions.
Classification of the family Ochotonidae
Five families are recognized in the order Lagomorpha: Mimolagidae
Erbajeva, 1986, Leporidae
Gray, 1821, Palaeolagidae
Dice, 1929, Prolagidae
Gureev, 1960, and Ochotonidae
Thomas, 1897. The family Ochotonidae includes two subfamilies, Sinolagomyinae
Gureev, 1960 and Ochotoninae
Thomas, 1897. The chronologic range of Sinolagomyinae is Oligocene to Miocene. Ochotoninae is known from the Miocene – Recent. The classification of genera (after
Erbajeva 1994, modified with new data from
Sen, 2003) is as follows:
Ochotonidae Thomas, 1897
Sinolagomyinae Gureev, 1960
Sinolagomys Bohlin, 1937
Austrolagomys Stromer, 1926
Kenyalagomys MacInnes, 1953
Oreolagus Dice, 1917
Bellatona Dawson, 1961
Heterolagus
Crusafont, Villata, and Truyols, 1955
Bellatonoides Sen, 2003
Ochotoninae Thomas, 1897
Marcuinomys Groizet, 1839
Lagopsis Schlosser, 1884
Albertona Lopez-Martinez, 1986
Alloptox Dawson, 1961
Paludotona Dawson, 1959
Proochotona Chomenko, 1914
Ochotonoides
Teilhard de Chardin and Young, 1931
Pliolagomys
Erbajeva, 1983
in Agadjanian and Erbajeva, 1983
Ochotonoma Sen, 1998
Ochotona Link, 1795
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