Buckman Hollow local fauna: Tables

author

TABLE 1. Measurements of the upper dentition of phenacodontids from the middle Clarkforkian (Cf2) Buckman Hollow local fauna, southwestern Wyoming.


Taxon	Specimen Number	P3L	P3W	P4L	P4W	M1L	M1W	M2L	M2W	M3L	M3W
Phenacodus vortmani	USNM 16693	7.9	7.3	—	—	—	—	—	—	—	—
	UCMP 173516	—	8.6	—	—	—	—	—	—	—	—
	USNM 21286 (R, except M3)	—	8.6	8.0	9.0	9.0	10.1	8.7	12.1	8.7	10.0
	USNM 16691 (L)	8.4	7.5	8.5	8.5	9.0	8.3	9.0	11.5	—	—
	USNM 16691 (R)	8.3	7.8	8.1	8.5	9.1	9.9	—	—	8.0	10.2
	USNM 406166	—	—	7.8	9.5	9.4	12.0	9.3	11.6	7.8	9.6
	USNM 20651	—	—	—	—	8.8	—	8.7	11.5	—	—
	USNM 16694	—	—	—	—	—	—	—	—	7.4	10.4
	USNM 20639	—	—	—	—	—	—	—	—	8.2	10.6
	USNM 406160	—	—	—	—	—	—	—	—	7.3	9.1
	USNM 406161	—	—	—	—	—	—	—	—	8.8	12.5
Phenacodus intermedius	USNM 406156	9.6	8.3	—	—	—	—	—	—	—	—
	UCMP 170704	10.2	9.3	9.8	11.3	—	—	—	—	—	—
	USNM 409474	—	—	11.0	11.8	—	—	—	—	—	—
	USNM 20649	—	—	9.9	10.9	—	—	11.0	14.0	—	—
Ectocion sp., cf. E. major	UCMP 173496	—	—	—	—	—	—	10.3	12.2	—	—
Ectocion osbornianus	USNM 406151	—	5.7	—	—	—	—	—	—	—	—
	USNM 20736 (L)	5.8	5.5	6.0	7.6	6.1	8.5	6.1	8.9	4.9	7.4
	USNM 20736 (R)	5.9	5.4	—	6.2	7.2	8.5	—	8.4	4.9	7.2
	USNM 406154	—	—	—	—	—	—	—	—	6.5	8.8

TABLE 2. Measurements of the lower dentition of phenacodontids from the middle Clarkforkian (Cf2) Buckman Hollow local fauna, southwestern Wyoming.


Taxon	Specimen Number	p3L	p3W	p4L	p4L	m1L	m1W	m2L	m2W	m3L	m3W
Phenacodus vortmani	USNM 406161	8.4	4.9	—	—	—	—	—	—	—	—
	USNM 20642	—	—	9.2	5.9	—	—	—	—	—	—
	USNM 20653	—	—	9.7	7.3	—	—	—	—	—	—
	USNM 20643	—	—	9.6	6.4	9.0	7.2	—	—	—	—
	UCMP 173495	—	—	—	—	9.0	7.3	—	—	—	—
	USNM 20654	—	—	—	—	9.0	7.9	9.9	8.2	—	—
	USNM 20638	—	—	—	—	9.4	8.7	9.5	7.8	10.0	6.7
	USNM 20650	—	—	—	—	9.5	7.9	9.8	8.0	—	7.4
	USNM 21286	—	—	—	—	—	—	9.3	—	9.4	—
	USNM 20637	—	—	—	—	—	—	—	—	10.8	7.7
	USNM 409469	—	—	—	—	—	—	—	—	10.4	6.9
Phenacodus intermedius	USNM 21287	12.0	7.7	12.6	10.0	12.5	11.4	13.2	12.7	—	—
	USNM 20644	—	—	11.8	7.9	12.1	9.8	—	—	—	—
	UCMP 170704	—	—	—	—	12.3	10.7	12.1	11.3	11.8	8.8
Ectocion osbornianus	USNM 20736 (R)	5.6	3.5	6.6	4.2	6.6	5.2	6.4	5.4	6.3	4.5
	USNM 20736 (L)	5.5	3.4	6.8	4.2	7.1	5.2	6.8	5.4	7.1	4.5
	USNM 16695	—	—	6.1	4.6	—	—	—	—	—	—
	USNM 20645	—	—	7.4	5.1	6.8	5.8	7.0	6.7	—	—
	USNM 20646	—	—	—	—	7.4	5.4	—	—	—	—
	UCMP 173493	—	—	—	—	—	—	6.7	5.9	—	—
	USNM 20648	—	—	—	—	6.7	5.3	6.6	5.7	—	—
	UCMP 173401	—	—	—	—	—	4.8	6.9	5.1	—	4.4
	USNM 20640	—	—	—	—	7.0	5.3	7.0	5.7	8.2	5.3
	USNM 406146	—	—	—	—	—	—	—	—	8.1	4.7

TABLE 3. Descriptive summary statistics of the upper dentition of phenacodontids from Buckman Hollow and other Clarkforkian and other earliest Wasatchian assemblages from the Western Interior of North America. Comparative data from Thewissen (1990), Burger (2012), and Rose et al. (2012).


Taxon	Sedimentary Basin		P3L	P3W	P4L	P4W	M1L	M1W	M2L	M2W	M3L	M3W
Phenacodus vortmani	Buckman Hollow	n	3	5	4	4	5	4	4	4	7	7
		Mean	8.2	8.0	8.1	8.9	9.1	10.1	8.9	11.7	8.0	10.4
		SD	0.2	0.6	0.3	0.5	0.2	1.6	0.3	0.3	0.6	1.1
		SE	0.1	0.3	0.1	0.2	0.1	0.8	0.2	0.1	0.2	0.4
		Minimum	7.9	7.3	7.8	8.5	8.8	8.3	8.7	11.5	7.3	9.1
		Maximum	8.4	8.6	8.5	9.5	9.4	12.0	9.3	12.1	8.8	12.5

	Bighorn Basin Cf 2	n				1	2	2	2	2	4	4
		Mean				9.6	9.4	11.2	9.8	10.5	9.5	8.9
		SD						0.6	0.4	0.4	0.1	0.7
		SE						0.4	0.3	0.3	0.1	0.4
		Minimum					9.0	10.9	9.5	10.4	8.6	7.7
		Maximum					9.8	11.4	10.0	10.6	10.3	9.6

Phenacodus intermedius	Buckman Hollow	n	2	2	3	3			1	1
		Mean	9.9	8.8	10.2	11.3			11.0	14.0
		SD	0.5	0.7	0.7	0.5
		SE	0.3	0.5	0.4	0.3
		Minimum	9.6	8.3	9.8	10.9
		Maximum	10.2	9.3	11.0	11.8

	Bighorn Basin Cf2	n	12	14	21	24	20	22	29	29	38	40
		Mean	10.7	9.9	10.7	11.5	12.3	13.9	13.2	14.3	11.7	11.0
		SD	0.7	0.7	0.7	0.9	0.9	1.1	1.0	0.9	0.9	1.0
		SE	0.2	0.2	0.1	0.2	0.2	0.2	0.2	0.2	0.1	0.2
		Minimum	9.5	8.9	9.2	9.0	10.7	11.8	10.9	12.4	10.2	8.8
		Maximum	12.1	11.4	11.6	13.1	14.5	16.3	16.1	15.7	14.0	14.2

	Bighorn Basin Wa0	n	1	1	1	1
		Mean	8.0	8.4	8.3	9.2
		SD
		SE
		Minimum
		Maximum

Ectocion sp., cf. E. major	Buckman Hollow	n							1	1
		Mean							10.3	12.2
		SD
		SE
		Minimum
		Maximum

Ectocion osbornianus	Buckman Hollow	n	2	3	1	2	2	2	1	2	3	3
		Mean	5.9	5.5	6.0	6.9	6.7	8.5	6.1	8.7	5.4	7.8
		SD	0.1	0.1		1.0	0.8	0.0		0.4	0.9	0.9
		SE	0.1	0.1		0.7	0.5	0.0		0.3	0.5	0.5
		Minimum	5.8	5.4		6.2	6.1	8.5		8.4	4.9	7.2
		Maximum	5.9	5.7		7.6	7.2	8.5		8.9	6.5	8.8

	Bighorn Basin Cf2	n	45	46	53	54	85	88	96	108	63	66
		Mean	6.2	6.2	6.6	7.8	7.2	8.7	7.6	8.9	6.2	7.0
		SD	0.3	0.5	0.4	0.6	0.4	0.5	0.5	0.5	0.5	0.5
		SE	0.1	0.1	0.1	0.1	0.0	0.1	0.1	0.1	0.1	0.1
		Minimum	5.2	4.9	5.8	5.6	6.4	7.3	6.2	7.7	5.2	5.6
		Maximum	6.7	6.9	7.2	8.8	8.0	10.0	8.9	10.1	7.5	8.0

Ectocion parvus	Bighorn Basin Cf2	n	2	2	2	2	4	4	8	7	5	5
		Mean	5.2	5.1	5.6	6.6	5.7	7.4	6.3	7.3	5.2	6.1
		SD	0.3	0.2	0.2	0.4	0.1	0.1	0.4	0.5	0.7	0.5
		SE	0.2	0.2	0.1	0.3	0.0	0.0	0.1	0.2	0.3	0.2
		Minimum	5.0	5.0	5.5	6.3	5.6	7.3	5.7	6.7	4.4	5.5
		Maximum	5.4	5.3	5.7	6.8	6.2	7.4	7.0	7.8	5.8	6.6

	Bighorn Basin Wa0	n	2	2	3	4	7	7	15	15	9	9
		Mean	4.7	4.7	5.4	6.5	5.6	6.8	5.6	7.2	4.9	6.2
		SD	0.3	0.6	0.2	0.4	0.3	0.6	0.3	0.5	0.8	0.8
		SE	0.2	0.4	0.1	0.2	0.1	0.2	0.1	0.1	0.3	0.3
		Minimum	4.6	4.3	5.2	6.0	5.3	6.1	4.9	6.4	4.1	4.9
		Maximum	4.9	5.1	5.5	6.9	6.0	7.5	6.2	7.9	6.6	7.2

	Piceance Creek	n	1	1	1	1	1	1	1	1	4	4
		Mean	5.4	4.9	5.4	6.1	5.8	7.5	5.6	7.5	4.9	7.0
		SD									0.1	0.1
		SE									0.1	0.1
		Minimum									4.9	6.8
		Maximum									5.2	7.1

TABLE 4. Descriptive summary statistics of the lower dentition of phenacodontids from Buckman Hollow and other Clarkforkian and other earliest Wasatchian assemblages from the Western Interior of North America. Comparative data from Thewissen (1990), Burger (2012), and Rose et al. (2012).


Taxon	Assemblage		p3L	p3W	p4L	p4W	m1L	m1W	m2L	m2W	m3L	m3W
Phenacodus vortmani	Buckman Hollow	n	1	1	3	3	5	5	4	3	4	4
		Mean	8.4	4.9	9.5	6.5	9.2	7.8	9.6	8.0	10.2	7.2
		SD			0.3	0.7	0.2	0.6	0.3	0.2	0.6	0.5
		SE			0.2	0.4	0.1	0.3	0.1	0.1	0.3	0.2
		Minimum			9.2	5.9	9.0	7.2	9.3	7.8	9.4	6.7
		Maximum			9.7	7.3	9.5	8.7	9.9	8.2	10.8	7.7

	Bighorn Basin Cf2	n	3	3	6	6	4	3	5	7	7	7
		Mean	8.5	5.6	9.5	6.5	9.6	7.7	9.7	8.1	9.8	6.9
		SD	0.4	0.2	0.3	0.5	0.5	0.4	0.2	0.3	0.8	0.6
		SE	0.2	0.1	0.1	0.2	0.2	0.2	0.1	0.1	0.3	0.2
		Minimum	8.1	5.4	9.1	5.7	9.0	7.2	9.5	7.7	8.5	6.3
		Maximum	8.8	5.7	9.8	7.1	10.0	7.9	10.0	8.4	11.0	7.6

Phenacodus intermedius	Buckman Hollow	n	1	1	2	2	3	3	2	2	1	1
		Mean	12.0	7.7	12.2	9.0	12.3	10.6	12.7	12.0	11.8	8.8
		SD			0.6	1.5	0.2	0.8	0.78	1.0
		SE			0.4	1.1	0.1	0.5	0.56	0.7
		Minimum			11.8	7.9	12.1	10.0	12.1	11.3
		Maximum			12.6	10.0	12.5	11.4	13.2	12.7

	Bighorn Basin Cf2	n	18	22	33	34	44	40	44	47	41	41
		Mean	10.9	6.9	11.9	8.6	12.4	10.3	12.8	11.3	13.1	9.8
		SD	0.6	0.6	0.8	0.7	0.7	0.6	0.7	0.8	0.9	0.6
		SE	0.1	0.1	0.1	0.1	0.1	0.1	0.1	0.1	0.1	0.1
		Minimum	10.2	5.4	10.2	7.5	11.1	9.3	11.2	9.6	11.4	8.4
		Maximum	12.1	8.1	13.5	10.1	14.0	11.6	14.3	12.9	15.2	10.7

	Bighorn Basin (Wa0)	n	2	2	2	2	2	1	1	1	2	2
		Mean	9.0	5.6	9.7	6.4	10.6	8.6	10.4	9.4	10.4	8.0
		SD	0.2	0.4	0.9	0.4	0.1				0.7	0.6
		SE	0.2	0.3	0.6	0.3	0.1				0.5	0.5
		Minimum	8.8	5.3	9.1	6.2	10.5				9.9	7.5
		Maximum	9.1	5.8	10.3	6.7	10.7				10.9	8.4

Ectocion osbornianus	Buckman Hollow	n	2	2	4	4	6	7	7	7	4	5
		Mean	5.6	3.4	6.7	4.5	6.9	5.3	6.8	5.7	7.4	4.7
		SD	0.1	0.1	0.5	0.4	0.3	0.3	0.2	0.5	0.9	0.4
		SE	0.1	0.1	0.3	0.2	0.1	0.1	0.1	0.2	0.5	0.2
		Minimum	5.5	3.4	6.1	4.2	6.6	4.8	6.4	5.4	6.3	4.4
		Maximum	5.6	3.5	7.4	5.1	7.4	5.9	7.0	6.7	8.2	5.3

	Bighorn Basin Cf2	n	49	52	98	103	173	181	178	187	91	105
		Mean	6.6	3.7	6.7	4.7	6.6	5.2	6.7	5.6	7.2	4.8
		SD	0.4	0.3	0.4	0.3	0.3	0.3	0.4	0.4	0.4	0.3
		SE	0.1	0.0	0.0	0.0	0.0	0.0	0.0	0.0	0.1	0.0
		Minimum	5.7	3.0	5.8	4.1	5.8	4.5	5.6	4.5	6.3	4.1
		Maximum	7.3	4.3	7.7	5.8	7.5	6.4	7.6	6.6	8.2	6.1

Ectocion parvus	Bighorn Basin Cf2	n	3	4	14	14	11	9	10	14	6	8
		Mean	5.8	3.1	5.8	3.8	5.5	4.4	5.5	4.6	5.9	3.8
		SD	0.3	0.2	0.3	0.2	0.2	0.3	0.4	0.3	0.4	0.3
		SE	0.2	0.1	0.1	0.1	0.1	0.1	0.1	0.1	0.2	0.1
		Minimum	5.5	2.8	5.1	3.4	5.2	3.8	4.6	4.2	5.1	3.4
		Maximum	6.0	3.2	6.3	4.2	5.9	4.8	5.9	5.2	6.2	4.3

	Bighorn Basin Wa0	n	8	8	15	15	19	18	26	27	19	18
		Mean	5.5	3.0	5.7	3.8	5.5	4.4	5.7	4.6	5.9	3.8
		SD	0.4	0.2	0.3	0.4	0.3	0.2	0.5	0.3	0.4	0.3
		SE	0.1	0.1	0.1	0.1	0.1	0.1	0.1	0.1	0.1	0.1
		Minimum	4.9	2.6	4.8	2.7	5.1	4.0	3.8	4.2	5.5	3.4
		Maximum	5.9	3.3	6.1	4.3	6.2	4.9	6.7	5.3	6.6	4.6

	Piceance Creek	n			1	1	1	1	1	1	3	3
		Mean			5.6	3.4	5.4	4.1	6.2	4.9	6.6	4.3
		SD									0.2	0.3
		SE									0.1	0.2
		Minimum									6.4	4.0
		Maximum									6.7	4.6

TABLE 5. Chart comparing current faunal lists for Clarkforkian faunas of southwestern Wyoming and northern Colorado. Solid circles indicate taxa referred to a fauna based on figured or published voucher specimens; open circles indicate taxa known only from faunal lists. Primary references for each locality are as follows: 1) Buckman Hollow – Gazin 1942, 1956; Dorr and Gingerich, 1980; Rose 1981, Gingerich 1976, Thewissen 1990; this paper; 2) Piceance Creek Clarkforkian composite – Patterson, 1933, 1934, 1937, 1939; Patterson and West, 1973; Kihm, 1984; Lucas, 1998; Burger and Honey, 2008; Burger, 2012; 3) Great Divide Basin – Anemone and Dirks, 2009; 4) Big Multi Quarry – Wilf et al., 1998; Dawson and Beard, 1996; Chester and Beard, 2012; 5) Togwotee Pass or lower variegated sequence Cf composite fauna in Purdy Basin – McKenna, 1980; Rose, 1981.


		Buckman Hollow	Piceance Creek Clarkforkian composite	Great Divide Basin	Big Multi	Purdy Basin Cf
Multituberculata
Eucosmodontidae	Neoliotomus conventus	●	○	—	○	—
Microcosmodontidae	Microcosmodon conus	—	—	—	●	—
Neoplagiaulacidae	Parectypodus sp.	—	—	○	●	—
Neoplagiaulacidae	Ectypodus sp.	—	—	—	●	—
Ptilodontidae	Prochetodon sp.	●	○	○	—	—
Marsupialia
Didelphidae	Peradectes sp.	—	—	○	○	—
Arctostylopida
Arctostylopidae	Arctostylops steini	—	—	—	—	○
"Condylarthra"
Arctocyonidae	Anacodon nexus	●	—	—	—	○
Arctocyonidae	Lambertocyon ischyrus	—	○	—	—
Arctocyonidae	Chriacus sp.	—	—	—	○	○
Oxyclaenidae	Thryptacodon sp.	○	○	○	—	—
Phenacodontidae	Ectocion osbornianus	●	○	○	○	●
Phenacodontidae	Ectocion parvus	—	●	—	—	○
Phenacodontidae	Ectocion major	cf	●	—	—	—
Phenacodontidae	Phenacodus intermedius	●	○	○	○	●
Phenacodontidae	Phenacodus vortmani	●	○	○	—	○
Apheliscidae	Haplomylus simpsoni	—	○	—	—	●
Apheliscidae	Apheliscus nitidus	●	—	○	○	●
Apheliscidae	Aletodon sp.	—	—	—	○	○
Creodonta
Oxyaenidae	Oxyaena transiens	—	—	—	—	○
Oxyaenidae	Oxyaena platypus	—	—	—	—	○
Oxyaenidae	Dipsalidictis krausei	●	—	—	—	—
Hyaenodontidae	cf. Prolimnocyon atavus	—	—	—	—	○
Hyaenodontidae	?Dipsalodon matthewi	—	—	—	—	●
Hyaenodontidae	Dipsalodon n. sp.	—	—	—	—	●
Dinocerata
Prodinoceratidae	Probathyopsis harrisorum	●	●	—	○	—
Mesonychia
Mesonychidae	Dissacus navajovius	●	○	—	—	○
Palaeanodonta
Epoicotheriidae	Palaeanodon cf. parvulus		—	—	○	—
Cimolesta
Barylambdidae	Barylambda faberi	●	●	—	—	—
Coryphodontidae	Coryphodon	—	—	—	—	○
Cyriacotheriidae	Cyriacotherium psamminum	—	—	—	○	—
Titanoideidae	Titanoides primaevus		○	—	—	—
Pantolesta
Pantolestidae	Palaeosinopa sp.	—	—	—	○	○
Proteutheria
Palaeoryctidae	Palaeoryctes cf. punctatus	—	—	—	○	—
Carnivora
Viverravidae	Protictis proteus	—	○	—	—	○
Viverravidae	Viverravus sp.	?	—	—	○	●
Viverravidae	Didymictis sp.	●	—	○	○	○
Dermoptera
	Planetetherium sp.	—	—	—	○	●
Primates
Microsyopidae	Arctodontomys sp.	—	—	—	○	—
Microsyopidae	Arctodontomys simplicidens	—	—	—	○	—
Plesiadapidae	Chiromyoides caesor	—	●	—	○	—
Plesiadapidae	Plesiadapis dubius	●	●	●	○	○
Plesiadapidae	Plesiadapis cookei	●	—	●	○	○
Plesiadapidae	cf. Plesiadapis	—	—	●	—	—
Paromomyidae	Phenacolemur cf. pagei	—	—	●	○	○
Paromomyidae	Phenacolemur cf. simonsi	—	—	—	○	—
Paromomyidae	cf. Ignacius sp.	—	—	—	○	—
Carpolestidae	Carpolestes nigridens	●	—	●	○	○
Micromomyidae	Dryomomys dulcifer	—	—	—	●	—
Micromomyidae	Tinimomys tribos	—	—	—	●	—
Lipotyphla
Erinaceidae	Diacocherus minutus	—	—	○	○	—
Erinaceidae	Leipsanolestes	—	—	—	○	●
Nyctitheriidae	Ceutholestes sp.	—	—	—	○	—
Nyctitheriidae	Limaconyssus sp.	—	—	—	○	—
Nyctitheriidae	Wyonycteris sp.	—	—	—	○	—
Nyctitheriidae	Plagioctenodon sp.	—	—	○	○	—
Rodentia
Ischyromyidae	Paramys adamus	●	—	●	○	—
Ischyromyidae	P. cf. P. atavus	—	—	—	—	○
Ischyromyidae	P. cf. P. excavatus	—	—	—	—	○
Alagomyidae	Alagomys russelli	—	—	—	●	—
Taeniodonta
Stylinodontidae	Ectoganus gliriformis	●	○	—	—	○
Tillodonta
Esthonychidae	Azygonyx xenicus	—	—	—	○	●

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Buckman Hollow local fauna: Tables