TABLE 1. Experimental parameters and associated burrow architectures. Terrarium sizes are in gallons. Sediment indicates the relative percentage of O = organic (coconut fiber), S = soil (clay loam), and Sa = fine- to medium-grained carbonate sand. Depth is the total depth of the sediment in centimeters. % Moisture is the mean moisture content of the sediment. Duration is split into two categories: expected time to completion of trial and the actual trial duration (in parentheses). Key to burrow architectures and modifications: V = vertical burrow, SV = subvertical burrow, H = helical burrow, J = J-shaped burrow, O = O-shaped burrow (T1=Type 1, T2=Type 2), c = chamber(s), b = branch(es), m = multiple entrances.
|
Experiment 1: Basic Morphology |
||||||||
|
Species |
Specimens |
Terrarium |
Sediment |
Depth |
% Moisture |
Duration |
Burrow #s |
Burrow Architectures |
|
N. americanus |
5 |
30 |
50/50 (O/S) |
20 |
37% |
7 (12) |
NA26 A-B |
SV, O (T1) |
|
N. americanus |
5 |
65 |
50/50 (O/S) |
45 |
32% |
7 (12) |
NA23 |
SV |
|
N. americanus |
1 |
30 |
50/50 (O/S) |
20 |
35% |
7 (8) |
No Burrow |
None |
|
N. americanus |
1 |
30 |
50/50 (O/S) |
20 |
30% |
7 (8) |
NA24 |
SVc |
|
N. americanus |
1 |
65 |
50/50 (O/S) |
45 |
40% |
7 (7) |
NA28 |
V |
|
N. americanus |
5 |
30 |
50/50 (O/S) |
20 |
35% |
14 (21) |
NA25 A-B |
SV, H |
|
N. americanus |
5 |
65 |
50/50 (O/S) |
43 |
37% |
14 (14) |
NA21 A-F |
SV, SVb, SVc, Hc |
|
N. americanus |
1 |
30 |
50/50 (O/S) |
20 |
35% |
14 (14) |
NA1 |
H |
|
N. americanus |
1 |
65 |
50/50 (O/S) |
45 |
35% |
14 (19) |
No Burrow |
None |
|
Experiment 2: Sediment Composition |
||||||||
|
Species |
Specimens |
Tank Size |
Sediment |
Depth |
% Moisture |
Duration |
Burrow #s |
Burrow Architectures |
|
F. penneri |
5 |
30 |
50/50 (Sa/S) |
16 |
50% |
14 (94) |
FP6 A-E |
SV, OC (T1) |
|
F. penneri |
5 |
65 |
100 (S) |
39 |
54% |
7 (8) |
FP7 A-H |
V, Vh, SV,SVb, SVcb, J |
|
F. penneri |
4 |
30 |
50/50 (Sa/S) |
19 |
70% |
14 (54) |
No Burrow |
None |
|
F. penneri |
5 |
30 |
25/75 (Sa/S) |
20 |
50% |
14 (14) |
FP5 A-G |
V, SV, SVc, O (T1), O (T2), Ob (T2) |
|
F. penneri |
1 |
30 |
50/50 (Sa/S) |
19 |
58% |
14 (19) |
FP4 A-B |
V, H |
|
F. penneri |
1 |
30 |
25/75 (Sa/S) |
20 |
52% |
14 (41) |
No Burrow |
None |
|
N. americanus |
5 |
30 |
25/75 (Sa/S) |
20 |
51% |
14 (23) |
NA32 |
SV |
|
N. americanus |
1 |
30 |
25/75 (Sa/S) |
20 |
60% |
14 (14) |
NA31 A-B |
O (T2), Om (T2) |
|
N. americanus |
1 |
65 |
100 (S) |
39 |
60% |
14 (51) |
NA33 A-C |
V, H |
|
Experiment 3: Sediment Moisture |
||||||||
|
Species |
Specimens |
Tank Size |
Sediment |
Depth |
% Moisture |
Duration |
Burrow #s |
Burrow Architectures |
|
F. penneri |
5 |
65 |
50/50 (O/S) |
45 |
49% |
14 (63) |
FP2 A-M |
V, SV, J, Jb |
|
F. penneri |
1 |
65 |
50/50 (O/S) |
45 |
50% |
14 (13) |
FP3 A-I |
V, SV |
|
F. penneri |
1 |
65 |
50/50 (O/S) |
45 |
50% |
14 (14) |
FP1 |
Hb |
|
N. americanus |
5 |
65 |
50/50 (O/S) |
48 |
50% |
14 (22) |
NA29 A-C |
V, H, O (T1) |
|
N. americanus |
1 |
65 |
50/50 (O/S) |
50 |
50% |
14 (21) |
NA30 A-F |
V, Vh, SV, SVc |
TABLE 2. Quantitative measurements of burrow casts produced by Narceus americanus.
|
NA32 |
NA23 |
NA25A |
NA21A |
NA21B |
NA21C |
NA30B |
NA30D |
NA21D |
NA24 |
NA21E |
NA26A |
NA30E |
|
|
Architecture |
SV |
SV |
SV |
SV |
SV |
SV |
SV |
SV |
SVb |
SVc |
SVc |
SVc |
SVc |
|
Surface Openings |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
|
Maximum Depth (cm) |
4.1 |
6.7 |
3.0 |
1.9 |
4.0 |
7.1 |
2.6 |
5.9 |
11.9 |
4.0 |
13.4 |
5.8 |
6.6 |
|
Total Length (cm) |
4.3 |
8.2 |
9.4 |
2.3 |
4.3 |
8.0 |
3.1 |
6.8 |
13.8 |
7.2 |
15.7 |
6.8 |
7.5 |
|
Maximum Width (mm) |
9.3 |
16.9 |
11.7 |
13.0 |
14.5 |
11.5 |
9.6 |
10.8 |
11.7 |
17.6 |
16.8 |
26.2 |
16.5 |
|
Minimum Width (mm) |
8.5 |
8.0 |
7.8 |
10.5 |
11.0 |
8.8 |
9.5 |
7.5 |
8.6 |
7.1 |
7.2 |
11.2 |
6.0 |
|
Average Width (mm) |
8.8 |
10.4 |
10.2 |
11.8 |
12.4 |
9.3 |
9.6 |
9.3 |
10.2 |
10.9 |
10.5 |
15.6 |
11.4 |
|
Maximum Height (mm) |
9.8 |
9.5 |
10.1 |
12.5 |
12.3 |
8.8 |
9.8 |
10.9 |
11.6 |
9.7 |
10.4 |
16.4 |
10.0 |
|
Minimum Height (mm) |
8.6 |
7.3 |
5.7 |
11.8 |
10.5 |
7.4 |
8.2 |
8.1 |
6.3 |
6.0 |
5.8 |
8.9 |
7.0 |
|
Average Height (mm) |
9.2 |
8.3 |
7.9 |
12.2 |
11.4 |
8.1 |
9.0 |
9.4 |
9.3 |
8.0 |
9.1 |
11.6 |
8.4 |
|
Average W/H Ratio |
1.0 |
1.3 |
1.3 |
1.0 |
1.1 |
1.1 |
1.1 |
1.0 |
1.1 |
1.3 |
1.2 |
1.4 |
1.3 |
|
Maximum Circumference (cm) |
3 |
4.1 |
3.7 |
4.2 |
4.3 |
3.1 |
2.9 |
3.5 |
3.6 |
4.6 |
4.3 |
8.4 |
4.7 |
|
Minimum Circumference (cm) |
2.7 |
2.7 |
2.5 |
3.5 |
3.3 |
2.2 |
2.8 |
2.7 |
2.6 |
2.5 |
2.3 |
3.3 |
2.5 |
|
Average Circumference (cm) |
2.8 |
3.1 |
3.1 |
3.9 |
3.7 |
2.7 |
2.9 |
3.1 |
3.1 |
3.3 |
3.2 |
4.9 |
3.5 |
|
Maximum Slope (Degrees) |
70 |
65 |
45 |
33 |
73 |
90 |
70 |
80 |
90 |
67 |
81 |
55 |
90 |
|
Minimum Slope (Degrees) |
70 |
50 |
25 |
33 |
70 |
25 |
70 |
60 |
40 |
20 |
55 |
55 |
65 |
|
Average Slope (Degrees) |
70 |
38 |
35 |
33 |
72 |
57 |
70 |
70 |
63 |
44 |
64 |
55 |
78 |
|
Branching Angle (Degrees) |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
60 |
n/a |
n/a |
n/a |
n/a |
|
Complexity |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
4.0 |
3.0 |
4.0 |
3.0 |
4.0 |
|
Tortuosity |
1.0 |
1.2 |
1.7 |
1.0 |
1.1 |
1.1 |
1.2 |
1.2 |
1.1 |
1.5 |
1.1 |
1.1 |
1.1 |
TABLE 2 (continued).
|
NA28 |
NA29C |
NA30A |
NA30C |
NA33A |
NA33B |
NA30F |
NA1 |
NA25B |
NA33C |
NA29B |
NA21F |
||
|
Architecture |
V |
V |
V |
V |
V |
V |
Vh |
H |
H |
H |
Hb |
Hc |
|
|
Surface Openings |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
|
|
Maximum Depth (cm) |
2.4 |
1.9 |
2.5 |
2.3 |
3.1 |
2.2 |
5.9 |
2.8 |
2.5 |
3.0 |
3.4 |
2.9 |
|
|
Total Length (cm) |
3.0 |
1.9 |
2.5 |
2.6 |
3.1 |
3.0 |
11.8 |
7.0 |
8.4 |
6.8 |
10.7 |
10.0 |
|
|
Maximum Width (mm) |
14.0 |
11.7 |
9.3 |
10.4 |
12.0 |
13.8 |
10.4 |
10.2 |
11.0 |
11.8 |
10.6 |
16.5 |
|
|
Minimum Width (mm) |
14.0 |
11.7 |
7.0 |
9.3 |
10.4 |
13.0 |
7.2 |
7.4 |
9.0 |
7.0 |
7.4 |
6.8 |
|
|
Average Width (mm) |
14.0 |
11.7 |
8.2 |
9.9 |
11.2 |
13.4 |
9.1 |
8.9 |
10.1 |
9.6 |
8.4 |
11.6 |
|
|
Maximum Height (mm) |
13.8 |
9.0 |
8.8 |
9.3 |
14.7 |
14.1 |
9.6 |
9.8 |
9.1 |
11.7 |
9.2 |
11.5 |
|
|
Minimum Height (mm) |
9.5 |
9.0 |
7.7 |
8.0 |
11.1 |
10.6 |
6.7 |
6.3 |
5.8 |
8.5 |
6.1 |
8.0 |
|
|
Average Height (mm) |
11.7 |
9.0 |
8.3 |
8.7 |
12.9 |
12.4 |
8.3 |
8.3 |
8.0 |
9.8 |
7.8 |
9.7 |
|
|
Average W/H Ratio |
1.2 |
1.3 |
1.0 |
1.1 |
0.9 |
1.1 |
1.1 |
1.1 |
1.3 |
1.0 |
1.1 |
1.2 |
|
|
Maximum Circumference (cm) |
5 |
3.5 |
2.8 |
3.1 |
4.3 |
4.4 |
3.2 |
3.2 |
3.3 |
4.3 |
3 |
4.3 |
|
|
Minimum Circumference (cm) |
3.7 |
3.5 |
2.4 |
2.8 |
3.5 |
3.9 |
2.1 |
2.2 |
3 |
2.3 |
2.5 |
2.3 |
|
|
Average Circumference (cm) |
4.4 |
3.5 |
2.6 |
3.0 |
3.9 |
4.2 |
2.8 |
2.8 |
3.2 |
3.1 |
2.7 |
3.4 |
|
|
Maximum Slope (Degrees) |
85 |
80 |
90 |
80 |
90 |
90 |
90 |
60 |
50 |
80 |
60 |
55 |
|
|
Minimum Slope (Degrees) |
85 |
80 |
90 |
80 |
90 |
90 |
0 |
15 |
0 |
50 |
60 |
0 |
|
|
Average Slope (Degrees) |
85 |
80 |
90 |
80 |
90 |
90 |
45 |
45 |
25 |
65 |
75 |
25 |
|
|
Branching Angle (Degrees) |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
90 |
n/a |
|
|
Complexity |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
4.0 |
4.0 |
|
|
Tortuosity |
1.3 |
1.0 |
1.0 |
1.1 |
1.0 |
1.0 |
2.0 |
3.5 |
5.6 |
2.3 |
1.5 |
1.2 |
TABLE 2 (continued).
|
NA29A |
NA26B |
NA31A |
NA31B |
Total Mean |
||
|
Architecture |
O (T1) |
O (T1) |
O (T2) |
O (T2) |
||
|
Surface Openings |
1 |
1 |
2 |
1 |
1 |
|
|
Maximum Depth (cm) |
3.4 |
6.9 |
2.5 |
1.9 |
4.4 |
|
|
Total Length (cm) |
10.6 |
12.5 |
12.3 |
10.0 |
7.4 |
|
|
Maximum Width (mm) |
12.9 |
18.0 |
13.4 |
10.9 |
13.2 |
|
|
Minimum Width (mm) |
8.3 |
7.4 |
8.7 |
9.1 |
8.8 |
|
|
Average Width (mm) |
10.5 |
10.9 |
10.4 |
10.1 |
10.6 |
|
|
Maximum Height (mm) |
9.6 |
11.3 |
12.7 |
11.7 |
11.0 |
|
|
Minimum Height (mm) |
6.0 |
6.7 |
9.3 |
8.9 |
7.9 |
|
|
Average Height (mm) |
8.2 |
9.3 |
10.4 |
9.9 |
9.5 |
|
|
Average W/H Ratio |
1.3 |
1.2 |
1.0 |
1.0 |
1.1 |
|
|
Maximum Circumference (cm) |
3.5 |
4.8 |
3.9 |
3.6 |
4.0 |
|
|
Minimum Circumference (cm) |
2.7 |
2.3 |
2.9 |
3.1 |
2.8 |
|
|
Average Circumference (cm) |
3.1 |
3.4 |
3.5 |
3.3 |
3.3 |
|
|
Maximum Slope (Degrees) |
60 |
60 |
90 |
20 |
71 |
|
|
Minimum Slope (Degrees) |
15 |
60 |
0 |
20 |
47 |
|
|
Average Slope (Degrees) |
38 |
60 |
43 |
20 |
59 |
|
|
Branching Angle (Degrees) |
n/a |
n/a |
n/a |
n/a |
75 |
|
|
Complexity |
3.0 |
3.0 |
6.0 |
2.0 |
2.6 |
|
|
Tortuosity |
43.0 |
39.0 |
1.8 |
99.0 |
7.6 |
TABLE 3. Quantitative measurements of burrow casts produced by Floridobolus penneri.
|
FP2C |
FP3C |
FP3D |
FP3E |
FP3F |
FP3G |
FP3H |
FP3I |
FP2E |
FP2F |
FP2G |
FP6A |
|
|
Architecture |
SV |
SV |
SV |
SV |
SV |
SV |
SV |
SV |
SV |
SV |
SV |
SV |
|
Surface Openings |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
|
Maximum Depth (cm) |
3.5 |
1.7 |
3.3 |
3.3 |
3.6 |
6.8 |
6.5 |
8.2 |
7.9 |
6.9 |
4.4 |
1.8 |
|
Total Length (cm) |
4.0 |
1.7 |
4.2 |
3.3 |
4.2 |
7.4 |
6.8 |
8.6 |
9.6 |
9.5 |
10.0 |
4.3 |
|
Maximum Width (mm) |
13.1 |
9.4 |
14.0 |
14.1 |
14.2 |
10.8 |
12.8 |
12.8 |
12.5 |
12.2 |
6.5 |
17.0 |
|
Minimum Width (mm) |
11.0 |
9.4 |
9.6 |
13.2 |
12.4 |
8.8 |
10.2 |
10.3 |
9.2 |
9.7 |
0.9 |
10.0 |
|
Average Width (mm) |
12.1 |
9.4 |
11.1 |
13.7 |
13.2 |
10.1 |
11.4 |
11.4 |
11.0 |
10.8 |
4.0 |
13.7 |
|
Maximum Height (mm) |
14.3 |
10.6 |
11.6 |
12.0 |
11.1 |
12.0 |
12.4 |
12.5 |
13.3 |
11.4 |
7.8 |
10.7 |
|
Minimum Height (mm) |
10.0 |
10.6 |
8.6 |
8.3 |
9.2 |
8.7 |
10.8 |
8.5 |
8.1 |
8.7 |
3.4 |
10.0 |
|
Average Height (mm) |
12.8 |
10.6 |
10.0 |
10.2 |
10.4 |
10.5 |
11.5 |
10.5 |
10.9 |
10.1 |
5.8 |
10.3 |
|
Average W/H Ratio |
1.0 |
0.9 |
1.1 |
1.4 |
1.3 |
1.0 |
1.0 |
1.1 |
1.0 |
1.1 |
0.7 |
1.3 |
|
Maximum Circumference (cm) |
4.4 |
3.2 |
4 |
4.4 |
4 |
3.7 |
4 |
4.1 |
3.9 |
3.7 |
3.7 |
4.5 |
|
Minimum Circumference (cm) |
3.4 |
3.2 |
3.1 |
3.7 |
3.8 |
2.9 |
3.1 |
3.2 |
3.2 |
2.9 |
2.4 |
2.8 |
|
Average Circumference (cm) |
4.0 |
3.2 |
3.4 |
4.1 |
3.9 |
3.3 |
3.6 |
3.6 |
3.6 |
3.4 |
3.2 |
3.7 |
|
Maximum Slope (Degrees) |
65 |
75 |
68 |
65 |
75 |
78 |
70 |
65 |
65 |
55 |
78 |
20 |
|
Minimum Slope (Degrees) |
65 |
75 |
68 |
65 |
75 |
78 |
70 |
65 |
65 |
55 |
78 |
20 |
|
Average Slope (Degrees) |
65 |
75 |
68 |
65 |
75 |
78 |
70 |
65 |
65 |
55 |
78 |
20 |
|
Branching Angle (Degrees) |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
|
Complexity |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
4.0 |
2.0 |
|
Tortuosity |
1.0 |
1.0 |
1.2 |
1.0 |
1.2 |
1.1 |
1.0 |
1.1 |
1.1 |
1.2 |
1.1 |
1.4 |
TABLE 3 (continued).
|
FP3A |
FP5B |
FP5C |
FP5D |
FP7A |
FP7C |
FP7E |
FP7F |
FP7H |
FP5E |
|
|
Architecture |
SV |
SV |
SV |
SV |
SV |
SV |
SV |
SVb |
SVcb |
SVc |
|
Surface Openings |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
|
Maximum Depth (cm) |
1.5 |
2.8 |
4.2 |
5.7 |
2.1 |
4.1 |
8.7 |
8.0 |
7.4 |
5.2 |
|
Total Length (cm) |
3.3 |
3.4 |
4.9 |
7.7 |
2.4 |
5.5 |
9.2 |
12.4 |
18.6 |
6.8 |
|
Maximum Width (mm) |
8.2 |
15.9 |
12.9 |
16.3 |
11.3 |
16.0 |
15.0 |
18.3 |
30.1 |
16.4 |
|
Minimum Width (mm) |
7.3 |
14.5 |
8.0 |
12.7 |
10.3 |
9.1 |
9.0 |
10.1 |
8.7 |
10.4 |
|
Average Width (mm) |
7.8 |
15.2 |
11.0 |
13.9 |
10.8 |
12.8 |
12.3 |
13.6 |
16.8 |
12.7 |
|
Maximum Height (mm) |
8.5 |
19.6 |
11.8 |
16.3 |
12.0 |
14.0 |
na |
12.8 |
19.0 |
24.7 |
|
Minimum Height (mm) |
7.6 |
13.3 |
10.9 |
11.8 |
10.8 |
10.7 |
na |
10.3 |
8.7 |
11.5 |
|
Average Height (mm) |
8.1 |
16.5 |
11.5 |
13.9 |
11.4 |
12.7 |
10.0 |
11.3 |
13.4 |
15.6 |
|
Average W/H Ratio |
1.0 |
1.0 |
1.0 |
1.0 |
0.9 |
0.9 |
1.2 |
1.2 |
1.3 |
0.9 |
|
Maximum Circumference (cm) |
2.7 |
5.7 |
4.1 |
5.2 |
3.6 |
4.5 |
3.9 |
4.5 |
8.1 |
6.2 |
|
Minimum Circumference (cm) |
2.6 |
4.6 |
3.2 |
3.3 |
3.1 |
3 |
2.9 |
3.1 |
2.6 |
3.3 |
|
Average Circumference (cm) |
2.7 |
5.2 |
3.7 |
4.5 |
3.4 |
3.9 |
3.6 |
4.0 |
5.0 |
4.7 |
|
Maximum Slope (Degrees) |
45 |
45 |
70 |
55 |
75 |
75 |
90 |
90 |
90 |
65 |
|
Minimum Slope (Degrees) |
45 |
45 |
70 |
55 |
75 |
50 |
65 |
50 |
20 |
45 |
|
Average Slope (Degrees) |
45 |
45 |
70 |
55 |
75 |
62 |
78 |
70 |
50 |
55 |
|
Branching Angle (Degrees) |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
20 |
55 |
n/a |
|
Complexity |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
3.0 |
4.0 |
3.0 |
|
Tortuosity |
1.6 |
1.1 |
1.1 |
1.2 |
1.1 |
1.3 |
1.1 |
1.1 |
1.5 |
1.3 |
TABLE 3 (continued).
|
FP3B |
FP2A |
FP2B |
FP2D |
FP4A |
FP5A |
FP7B |
FP2M |
FP7D |
|
|
Architecture |
V |
V |
V |
V |
V |
V |
V |
V |
Vh |
|
Surface Openings |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
|
Maximum Depth (cm) |
2.2 |
4.1 |
3.1 |
6.0 |
3.3 |
2.1 |
3.6 |
3.6 |
6.9 |
|
Total Length (cm) |
2.2 |
4.1 |
3.1 |
6.7 |
3.6 |
2.1 |
4.7 |
3.6 |
8.4 |
|
Maximum Width (mm) |
12.6 |
10.6 |
15.0 |
15.6 |
16.5 |
16.9 |
14.5 |
23.3 |
14.4 |
|
Minimum Width (mm) |
9.8 |
9.2 |
13.6 |
8.6 |
12.3 |
10.3 |
7.6 |
12.1 |
10.3 |
|
Average Width (mm) |
11.2 |
9.9 |
14.3 |
13.2 |
14.7 |
13.6 |
12.0 |
16.6 |
12.1 |
|
Maximum Height (mm) |
9.7 |
11.9 |
15.8 |
13.3 |
13.4 |
15.4 |
13.7 |
23.0 |
11.3 |
|
Minimum Height (mm) |
9.1 |
10.0 |
14.0 |
9.6 |
10.5 |
12.0 |
11.6 |
12.7 |
9.8 |
|
Average Height (mm) |
9.4 |
11.1 |
14.9 |
11.5 |
12.2 |
13.7 |
12.7 |
16.7 |
11.0 |
|
Average W/H Ratio |
1.2 |
0.9 |
1.0 |
1.1 |
1.2 |
1.0 |
0.7 |
1.0 |
1.1 |
|
Maximum Circumference (cm) |
3.6 |
3.8 |
5.2 |
5.6 |
5.5 |
4.6 |
4 |
6.5 |
3.8 |
|
Minimum Circumference (cm) |
3 |
3.4 |
4.6 |
2.6 |
3.5 |
3.4 |
3.2 |
3.8 |
3.3 |
|
Average Circumference (cm) |
3.3 |
3.6 |
4.9 |
4.4 |
4.6 |
4.0 |
3.6 |
5.2 |
3.6 |
|
Maximum Slope (Degrees) |
90 |
85 |
90 |
85 |
90 |
90 |
90 |
90 |
90 |
|
Minimum Slope (Degrees) |
90 |
85 |
90 |
60 |
90 |
90 |
40 |
90 |
55 |
|
Average Slope (Degrees) |
90 |
85 |
90 |
73 |
90 |
90 |
65 |
90 |
73 |
|
Branching Angle (Degrees) |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
n/a |
|
Complexity |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
|
Tortuosity |
1.0 |
1.0 |
1.0 |
1.1 |
1.1 |
1.0 |
1.2 |
1.0 |
1.2 |
TABLE 3 (continued).
|
FP2I |
FP2J |
FP2K |
FP2L |
FP7G |
FP2H |
FP4B |
FP1 |
FP5F |
FP6B |
FP5G |
Total Mean |
||||
|
Architecture |
J |
J |
J |
J |
J |
Jb |
H |
Hb |
Ob (T1) |
Oc (T1) |
Ob (T2) |
||||
|
Surface Openings |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
1 |
|||
|
Maximum Depth (cm) |
6.5 |
7.8 |
8.1 |
2.9 |
8.4 |
5.8 |
3.1 |
2.8 |
7.7 |
4.4 |
1.1 |
4.8 |
|||
|
Total Length (cm) |
10.7 |
11.9 |
11.8 |
7.5 |
14.7 |
13.0 |
11.1 |
11.1 |
17.5 |
13.0 |
10.1 |
7.6 |
|||
|
Maximum Width (mm) |
11.5 |
16.6 |
12.4 |
11.4 |
13.0 |
13.0 |
13.6 |
13.5 |
12.5 |
24.8 |
11.9 |
14.4 |
|||
|
Minimum Width (mm) |
8.4 |
9.3 |
9.5 |
7.9 |
9.0 |
7.2 |
10.0 |
7.5 |
9.1 |
10.0 |
9.0 |
9.7 |
|||
|
Average Width (mm) |
9.9 |
12.0 |
11.0 |
9.4 |
11.3 |
10.8 |
11.6 |
9.8 |
10.8 |
17.7 |
10.2 |
11.9 |
|||
|
Maximum Height (mm) |
13.0 |
16.5 |
14.8 |
12.1 |
15.7 |
14.7 |
11.4 |
7.0 |
13.7 |
16.0 |
11.2 |
13.5 |
|||
|
Minimum Height (mm) |
9.2 |
9.2 |
10.3 |
7.4 |
10.4 |
7.7 |
7.5 |
5.1 |
10.0 |
7.7 |
6.2 |
9.5 |
|||
|
Average Height (mm) |
10.7 |
11.8 |
13.0 |
9.5 |
12.4 |
11.0 |
10.0 |
6.1 |
12.2 |
12.1 |
8.3 |
11.4 |
|||
|
Average W/H Ratio |
0.9 |
1.0 |
0.9 |
1.0 |
0.9 |
1.0 |
1.2 |
1.6 |
0.9 |
1.5 |
1.3 |
1.1 |
|||
|
Maximum Circumference (cm) |
3.7 |
5.3 |
4.2 |
3.5 |
4.2 |
6.4 |
4.3 |
3.4 |
4.1 |
6.1 |
3.5 |
4.5 |
|||
|
Minimum Circumference (cm) |
2.9 |
2.8 |
3 |
2.6 |
3.1 |
2.4 |
3.1 |
2.2 |
3.2 |
2.6 |
2.7 |
3.1 |
|||
|
Average Circumference (cm) |
3.2 |
3.8 |
3.8 |
3.1 |
3.9 |
3.7 |
3.6 |
2.7 |
3.7 |
4.4 |
3.1 |
3.8 |
|||
|
Maximum Slope (Degrees) |
80 |
90 |
55 |
55 |
75 |
65 |
70 |
50 |
85 |
90 |
20 |
72 |
|||
|
Minimum Slope (Degrees) |
15 |
50 |
50 |
45 |
75 |
25 |
0 |
25 |
85 |
40 |
20 |
58 |
|||
|
Average Slope (Degrees) |
47 |
70 |
53 |
50 |
75 |
45 |
35 |
38 |
85 |
65 |
20 |
65 |
|||
|
Branching Angle (Degrees) |
n/a |
n/a |
n/a |
n/a |
n/a |
70 |
n/a |
10,45 |
12, 25 |
n/a |
10 |
31 |
|||
|
Complexity |
2.0 |
2.0 |
2.0 |
2.0 |
2.0 |
3.0 |
2.0 |
6.0 |
6.0 |
4.0 |
4.0 |
2.5 |
|||
|
Tortuosity |
2.2 |
2.0 |
2.0 |
5.8 |
2.4 |
1.4 |
4.3 |
1.1 |
17.6 |
53.0 |
1.4 |
3.1 |
TABLE 4. Rs values of the Spearman's rank correlation tests between sediment clay and moisture content and quantitative burrow properties. The number of burrow openings rarely varied between burrow casts and were, therefore, excluded from the analysis.
|
Narceus americanus |
||||||||
|
Length |
Depth |
Width |
Height |
Circum. |
Slope |
Complexity |
Tortuosity |
|
|
Clay |
-0.31 |
-0.10 |
0.01 |
0.42 |
-0.13 |
-0.39 |
-0.28 |
-0.30 |
|
Moisture |
-0.22 |
-0.33 |
-0.13 |
0.41 |
0.04 |
0.46 |
-0.10 |
-0.11 |
|
Floridobolus penneri |
||||||||
|
Length |
Depth |
Width |
Height |
Circum. |
Slope |
Complexity |
Tortuosity |
|
|
Clay |
0.15 |
0.19 |
0.24 |
0.36 |
0.22 |
0.03 |
-0.17 |
0.14 |
|
Moisture |
0.03 |
-0.07 |
0.26 |
-0.01 |
0.04 |
-0.01 |
0.09 |
0.11 |
