TABLE 1. Meristic counts of all studied specimens of †Paralates . Numbers not in bold are subject to some uncertainty.
Oligocene of Rouffach, no. Ruf | Upper Eocene of Isle of Wight (Chapelcorner Fish Bed) |
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Specimen # | 6a/b | 9_1 | 9_2 | 11/13_2 | 15a/b | 18 | P 59784 | P 59785 | P 59786 | C 23632 | C 23633 |
First dorsal fin (D1) | - | - | >V | - | VII | VII | VII | VII | VII | - | - |
Second dorsal fin (D2) | I,10 | I,9 | I,9 | I,9 | I,8 | I,10 | I,9 | I,10 | >7 | - | - |
Anal fin (A) | 10 | >8 | 10 | >7 | I,8 | I,8 | I,8 | I,9 | - | - | - |
C (branched, segmented) | 12 | 13 | ? | 13 | 12 | 12 | 12 | 13 | - | - | - |
C (unbranched, segmented) | 2 + 1 or 2 | 2 + 2 | |||||||||
C (unbranched, unsegmented) | 7 | 10 + ? | 10 + 12 | ||||||||
Ventral fin (V) | - | - | - | - | I,5 | - | I,5 | I,5 | - | - | - |
Pectoral fin (P) | - | >11 | 14 | >9 | 12 | - | >14 | 16 | >11 | >13 | >9 |
Vertebrae [postabdominal vert.] | >23 [17] | >21 [16] | >27 | >23 [17] | 29 [17] | 30 [18] | 30 [18] | 31 [19] |
>23 | >11 | 29 [17] |
Pterygiophore formula D1 | - | - | - | - | - | - | - | 3-2121100 | - | - | - |
TABLE 2. Morphometrics of the genus †Paralates in mm. Values standardized to standard length are given in brackets. Abbreviations: A = anal fin, C = caudal fin, D1 = first dorsal fin, D2 = second dorsal fin, D2C = distance between end of D2 and first procurrent ray of C, P = pectoral fin, V = pelvic fin.
# | Ruf. 11/13_2 | Ruf. 15a/b | Ruf. 18 | P 59784 | P 59785 | P 59786 |
Total length | 29.1 (115%) | 29.5 (118%) | 34.5 (119%) | 35.3 (116%) | 22.5 (116%) | >22.3 |
Standard length | 25.2 | 25.1 | 29.1 | 30.4 | 19.4 | >22.3 |
Max body height | 5.4 (21.4%) | 5.4 (21.5%) | 5.5 (18.9%) | 5.9 (19.4%) | 4.4 (22.7%) | - |
Head length | 9.4 (37.3%) | 6.9 (27.5%) | 7.1 (24.4%) | 9.5 (31.3%) | 5.6 (28.9%) | 6.6 |
Head height | 6.6 (26.1%) | 5.4 (21.5%) | 5.2 (17.9%) | 6.9 (22.7%) | 4.9 (25.3%) | - |
Horizontal eye diameter | 1.5 (5.9%) | 1.2 (4.8%) | 1.3 (4.5%) | - | 1.7 (8.8%) | 1.4 |
Dist snout to D1 | 11.7 (46.4%) | 9.2 (36.7%) | 10.8 (37.1%) | 11.7 (38.5%) | 7.8 (40.2%) | 7.9 |
Dist snout to D2 | 14.6 (57.9%) | 14.5 (57.8%) | 16.2 (55.7%) | 17.4 (57.2%) | 11.2 (57.7%) | 12.8 |
Dist snout to A | 15.8 (62.7%) | 15.7 (62.5%) | 16.5 (56.7%) | 18.1 (59.5%) | 11.5 (59,3%) | 13.3 |
Dist snout to P | 9.4 (37,3%) | 8.7 (34.7%) | - | 10.7 (35.2%) | 6.8 (35.1%) | 7.4 |
Dist snout to V | 9.2 (36.5%) | 8.2 (32.7%) | - | 10.6 (34.9%) | 6.6 (34.0%) | - |
Length D1 | >2.1 | >2.4 | >2.4 | >3.4 | >1.7 | - |
Length D2 | 2.8 (11.1%) | >2.4 | 4.4 (15.1%) | >4.3 | >3.1 | - |
Length A | 2.6 (10.3%) | >2.4 | >2.3 | >3.5 | >2.9 | - |
Length P | >0.8 | >1.3 | - | >1.9 | >0.8 | >2.8 |
Length V | >1.2 | >1.5 | - | >2.7 | 2.7 (13.9%) | - |
Base length D1 | >0.6 | 2.3 (9.2%) | 2.3 (7.9%) | 2.6 (8.6%) | 1.6 (8.2%) | - |
Base length D2 | 3.1 (12.3%) | 3.0 (12.0%) | 4.0 (13.7%) | >3.5 | 2.4 (12.4%) | - |
Base length A | 3.0 (11.9%) | 2.5 (19.0%) | 3.7 (12.7%) | 3.6 (11.8%) | 1.9 (9.8%) | - |
Length caudal peduncle | 6.3 (25.0%) | 6.8 (27.1%) | 8.9 (30.6%) | 8.3 (27.3%) | 5.7 (29.4%) | - |
D2C | 6.7 (26.6%) | 7.3 (29.1%) | 8.4 (28.6%) | 7.5 (24.7%) | 5.0 (25.8%) | - |
Min height of caud ped | 2.0 (7.9%) | 1.7 (6.8%) | 2.6 (8.9%) | 3.0( 9.7%) | 2.2 (11.3%) | - |
Length of caudal fin | 3.8 (15.1%) | >3.8 | >5.0 | 5.4 (17.8%) | 3.6 (18.6%) | - |
Length of vertebrae | 0.5-0.6 (1.9-2.3%) | 0.6-0.7 (2.4-2.8%) | 0.6-0.8 (2.1-2.7%) | 0.5-0.7 (1.6-2.3%) | 0.4-0.5 (2.1-2.6%) | 0.4-0.5 |
TABLE 3. Meristic characters of recent gobioid taxa for which the pterygiophore formula 3-212110(0) has been observed in at least one species of the genus. The number of caudal fin rays refers to all segmented rays (branched and unbranched). Abbreviations: A = anal fin, C = caudal fin, D1 = first dorsal fin, D2 = second dorsal fin, P = pectoral fin, V = pelvic fin.
Taxon | D1 | D2 | A | C | V | P | Vertebrae [postabd] | Source | Family |
†Paralates bleicheri Sauvage, 1883 | VII | I,10 | I,9 | >12-13 | I,5 | 14 | 29 [17-18?] |
this study | inc. sed. |
†Paralates chapelcorneri n. sp. | VII | I,10 | I,8-9 | 14-17 | I,5 | 14-16 | 30?-31 [18?-19] | this study | inc. sed. |
Gobiodon okinawae Sawada, Arai & Abe, 1972 | VI | I,10 | I,9 | 15 | - | - | 27 [17] |
unpublished data | Gobiidae |
Gobiosoma hemigymnum (Eigenmann and Eigenmann 1888) | VII | I,11 | I,9 | 17 | I,5 | 17-19 | 27 [16] |
van Tassell et al. 2015 | Gobiidae |
Gobiosoma spp. | - | - | - | - | - | - | 27-28 [15-17] |
Birdsong et al. 1988 | Gobiidae |
Gobulus spp. | - | - | - | - | - | - | 27-28 [16-17] |
Birdsong et al. 1988 | Gobiidae |
Gobulus spp. | VII-VIII | I,9-13 | I,7-13 | 15-17 | I,5 | 15-18 | 27-28 [16-17] |
Hoese and Reader 2001 | Gobiidae |
Gobiomorphus alpinus Stokell, 1962 | III-VII | I,7-9 | I,8-10 | - | - | 16-19 | 29-31 | McDowall 1994 | Eleotridae |
Gobiomorphus spp. | III-VIII | I,7-10 | I,7-10 | - | - | 13-21 | 27-31 | McDowall 1975 | Eleotridae |
Gobiomorphus spp. | - | - | - | - | - | - | 28-31 [16-19] |
Birdsong et al. 1988 | Eleotridae |
Thalasseleotris | 16 | 27 [17] |
Hoese and Larson 1987 | Thalasseleotrididae |