TABLE 1. Species of Plesiosminthus described from the European Oligocene and Miocene.
Species | Type-locality |
P. schaubi Viret, 1926 (type-species) | Coderet, France |
P. myarion Schaub, 1930 | Chavroches, France |
P. promyarion Schaub, 1930 | Puy-de-Montdoury, F rance |
P. bavaricus Freudenberg, 1941 | Gaimersheim, Germany |
P. winistoerferi Engesser, 1987 | Brochene Fluh 53, Switzerland |
P. conjunctus Ziegler, 1994 | Herrlingen 8, Germany |
P. moralesi Álvarez Sierra, Daams and Lacomba Andueza, 1996 | Sayatón 1, Spain |
P. admyarion Comte, 2000 | Thézels, France |
P. meridionalis Comte, 2000 | Venelles inf., France |
P. margaritae n. sp. (this paper) | Mirambueno 1, Spain |
P. moniqueae n. sp. (this paper) | Pech Desse, France |
Plesiosminthus n. sp., Ziegler, 1994 | Herrlingen 8, Germany |
Plesiosminthus sp., Engesser and Mödden, 1987 | Mümliswil-Hardberg, Switzerland |
Plesiosminthus sp., Comte, 2000 | Venelles inf., France |
TABLE 2. Locality codes, species contents and most important reference of the localities studied. Zones are European mammal zones (Biochrom’97, 1997).
Code | Locality | Zone | Species | Reference |
BF53 | Brochene Fluh 53 | MP30 | P. winistoerferi | Engesser (1987) |
CANAL | Canales | MP28 | Plesiosminthus sp. | Alvarez Sierra et al. (1999) |
CHAVR | Chavroches | MN2 | P. myarion | Hugueney and Vianey-Liaud (1980) |
CLUZEL | Cluzel | MN2 | P. myarion | Hugueney and Vianey-Liaud (1980) |
COD3 | Coderet 3 | MP30 | P. schaubi | Hugueney (1969) |
DIEU | Dieupentale | MP30 | P. schaubi | Baudelot and Olivier (1978) |
FORN11 | Fornant 11 | MN1 | P. myarion | Engesser (1987) |
GAIM | Gaimersheim | MP28 | P. bavaricus | Kristkoiz (1992) |
HERR8 | Herrlingen 8 | MP28 | P. conjunctus | Ziegler (1994) |
LAUT2 | Lautern 2 | MN1 | P. myarion | Ziegler and Werner (1994) |
MIR1 | Mirambueno 1 | MP27 | P. margaritae | this paper |
MIR2A | Mirambueno 2A | MP27 | P. aff. conjunctus | this paper |
OBL | Oberleichtersbach | MP30 | P. winistoerferi | Engesser and Storch (2008) |
PARR | Parrales | MP29 | P. schaubi | Alvarez Sierra et al. (1999) |
PDES | Pech Desse | MP28 | P. promyarion | Hugueney and Vianey-Liaud (1980) |
PDES | Pech Desse | MP28 | P. moniqueae | this paper |
Pech-du-Fraysse | MP28 | P. promyarion | Hugueney and Vianey-Liaud (1980) | |
RDUBEY | Ruisseau du Bey | MP28 | P. promyarion | Engesser (1987) |
SAULC | Saulcet | MN1 | P. myarion | Hugueney and Vianey-Liaud (1980) |
SAY1 | Sayatón 1 | MP29 | P. moralesi | Alvarez et al. (1996) |
THZ | Thézels | MP30 | P. admyarion | Comte (2000) |
VENINF | Venelles inf. | MP30 | P. meridionalis | Comte (2000) |
VIV1 | Vivel del Río 1 | MP28 | P. cf. margaritae | this paper |
TABLE 3. Frequencies of character states of the protolophule of M2, number of observations, and morphology values. Ant. = anterior, trans = transverse, ant.plus = anterior plus an incomplete posterior connection. * protolophule posterior, not double.
Sample | ant. or trans | ant. plus | double | n | MV |
CHAVR | 0 | 3 | 97 | 33 | 0.985 |
FORN11 | |||||
VENINF | 4 | 0 | 96* | 25 | 0.960 |
THZ | 95 | 0 | 5 | 97 | 0.050 |
COD3 | 97 | 0 | 3 | 63 | 0.032 |
HERR8 | 17 | 8 | 75 | 12 | 0.790 |
GAIM | 39 | 52 | 9 | 23 | 0.350 |
PDES | 74 | 13 | 13 | 16 | 0.195 |
RDUBEY | |||||
VIV1 | 57 | 4 | 39 | 69 | 0.410 |
MIR2A | 19 | 7 | 74 | 27 | 0.775 |
MIR1 | 28 | 37 | 35 | 43 | 0.535 |
TABLE 4. P-values of t-test comparing the samples from THZ and COD3; df = degrees of freedom.
Molar | Comte (2000) |
this paper | df |
m1 | 3.18 | 3.092 | 205 |
m2 | 2.02 | -2.162 | 180 |
m3 | 3.00 | 2.828 | 127 |
M1 | 2.39 | -2.483 | 188 |
M2 | 2.06 | -2.178 | 172 |
M3 | 3.00 | 2.832 | 98 |
TABLE 5. Frequencies of character states of the protoconid hind arm of m2, number of observations, and morphology values. Data from Kristkoiz (1992, figure 84) and this paper.
Sample | complete | incomplete | absent | n | MV |
CHAVR | 12 | 50 | 38 | 50 | 0.630 |
FORN11 | 26 | 52 | 22 | 27 | 0.480 |
VENINF | 0 | 0 | 100 | 26 | 1.000 |
COD3 | 0 | 0 | 100 | 59 | 1.000 |
THZ | 9 | 9 | 82 | 127 | 0.865 |
HERR8 | 82 | 18 | 0 | 17 | 0.090 |
36 | 53 | 11 | 28 | 0.375 | |
GAIM | 50 | 41 | 9 | 31 | 0.295 |
PDES | 32 | 45 | 23 | 53 | 0.455 |
RDUBEY | 31 | 69 | 0 | 13 | 0.345 |
VIV1 | 31 | 16 | 53 | 52 | 0.610 |
MIR2A | 83 | 0 | 17 | 16 | 0.170 |
MIR1 | 60 | 26 | 14 | 41 | 0.270 |
TABLE 6. Frequencies of character states of the protoconid hind arm of m3, number of observations, and morphology values.
Sample | complete | incomplete | absent | n | MV |
CHAVR | 5.4 | 0.0 | 94.6 | 37 | 0.946 |
VENINF | 0.0 | 0.0 | 100.0 | 8 | 1.000 |
COD3 | 0.0 | 0.0 | 100.0 | 32 | 1.000 |
THZ | 0.0 | 0.0 | 100.0 | 97 | 1.000 |
HERR8 | 0.0 | 0.0 | 100.0 | 14 | 1.000 |
8.0 | 32.0 | 60.0 | 25 | 0.760 | |
GAIM | 14.9 | 27.3 | 63.6 | 11 | 0.730 |
PDES | 25.0 | 43.8 | 31.2 | 16 | 0.531 |
RDUBEY | 0.0 | 0.0 | 100.0 | 6 | 1.000 |
VIV1 | 5.1 | 0.0 | 94.9 | 39 | 0.949 |
MIR2A | 20.0 | 15.0 | 65.0 | 20 | 0.725 |
MIR1 | 23.2 | 11.6 | 65.1 | 43 | 0.710 |
TABLE 7. Number of specimens for samples in Figure 13.
Sample | m1 | m2 | m3 | M1 | M2 | M3 |
BF53 | 4 | 4 | 5 | 6 | 5 | 5 |
CHAVR | 36 | 48 | 39 | 49 | 33 | 15 |
COD3 | 65 | 59 | 32 | 71 | 63 | 13 |
GAIM | 34 | 32 | 11 | 28 | 23 | 3 |
HERR8 | 8 | 15 | 14 | 18 | 12 | 11 |
MIR1 | 17 | 41 | 45 | 49 | 43 | 37 |
PDES | 57 | 54 | 16 | 35 | 16 | 3 |
SAY1 | 7 | 6 | 1 | 3 | 3 | 1 |
THZ | 142 | 123 | 97 | 119 | 111 | 87 |
VENINF | 19 | 25 | 8 | 27 | 26 | 1 |