TABLE 1. Description of the Dubrovino section.
Layer No. | Elevation (m) |
Thickness (m) | Description |
8 | 12.7-13.0 | 0.3 | Modern soil |
7 | 12.0-12.7 | 0.7 | Sandy loam consistent, dry, whitish, with ferruginous interlayer in lower part |
6 | 7.1-12.0 | 4.9 | Horizontal alternation of medium and coarse-grained sands |
5 | 6.9-7.1 | 0.2 | Interlayer with fine and compact light-grey sand |
4 | 4.4-6.9 | 2.5 | Alternation of medium sands and coarse-grained sands; in right side coarse-grained sands are turning into sandy gravel with fragments fragment of cbivalve shells |
3 | 0.4-4.4 | 4.0 | Alternation of fine-grained sands, loamy sands and grey-brown loams |
2* | 0.1-0.4 | 0.3 | Grey loam with a streak of alluvial detritus |
1 | 0-0.1 | >0.1 | Rufous clay extending under water surface |
*Sample upper 0.2 m (12.6-12.8 m from the surface) with insect remains and phytodetritus for dating.
TABLE 2. Species composition and number of insect fragments in Dubrovino taphocoenosis. HD = head, PR = pronotum, EL = elytra, OT = other fragments, Nmin = minimum number of specimens.
No. | Species | Distribution* | Type of fragment | Nmin | |||
HD | PR | EL | OT | ||||
Carabidae | |||||||
1 | Notiophilus cf. aquaticus (Linnaeus, 1758) | + | - | - | 2 | - | 2 |
2 | Notiophilus fasciatus Mäklin, 1855 | North | - | - | 1 | - | 1 |
- | Notiophilus sp. | - | - | 1 | - | 1 | |
3 | Nebria cf. rubrofemorata Shilenkov, 1975 | East | - | - | 1 | - | 1 |
4 | Nebria subdilatata Motschulsky, 1844 | East | - | - | 1 | - | 1 |
5 | Diacheila polita (Faldermann, 1835) | North | - | - | 1 | - | 1 |
6 | Elaphrus cf. riparius (Linnaeus, 1758) | + | 1 | 2 | - | - | 2 |
- | Elaphrus sp. | - | - | - | 2 | 1 | 1 |
7 | Clivina fossor (Linnaeus, 1758) | + | - | 2 | 2 | - | 2 |
8 | Trechus cf. compactulus Belousov and Kabak, 1996 | East | - | - | 2 | - | 2 |
9 | Trechus sp. 1 | - | - | - | 3 | - | 2 |
10 | Bembidion (Plataphodes) cf. fellmanni (Mannerheim, 1823) | North | - | - | 1 | - | 1 |
- | Bembidion (Plataphodes) sp. | - | - | - | 1 | - | 1 |
11 | Bembidion (Plataphus) sp. | - | - | - | 2 | - | 2 |
12 | Bembidion (Peryphus) dauricum (Motschulsky, 1844) | East | - | - | 4 | - | 3 |
13 | Bembidion (Asioperyphus) spp. | - | 4 | - | - | - | 4 |
14 | Bembidion (Pamirium) sp. | South | - | - | 1 | - | 1 |
- | Bembidion sp. | - | - | - | 1 | 4 | 1 |
15 | Pogonus iridipennis Nicolai, 1822 | South | - | 1 | - | - | 1 |
16 | Pogonus punctulatus Dejean, 1828 | South | - | 3 | 3 | - | 3 |
17 | Patrobus cf. septentrionis Dejean, 1828 | + | - | - | 1 | - | 1 |
18 | Poecilus (Derus) cf. ravus (Lutshnik, 1922) | East | - | 6 | 9 | - | 5 |
19 | Poecilus (Derus) sp. 1 | - | - | 12 | 6 | - | 11 |
20 | Poecilus (Poecilus) fortipes (Chaudoir, 1850) | + | - | 1 | - | - | 1 |
21 | Pterostichus (Lyperopherus) mirus (Tschitschérine, 1894) | East | - | 1 | - | - | 1 |
30 | Pterostichus (Adelosia) macer (Marsham, 1802) | South | - | 1 | - | - | 1 |
22 | Pterostichus (Cryobius) cf. brevicornis (Kirby, 1837) | North | - | 2 | 1 | - | 2 |
23 | Pterostichus (Cryobius) spp. | North? | - | 3 | 6 | - | 4 |
24 | Pterostichus (Petrophilus) cf. tomensis (Gebler, 1847) | East | - | 2 | - | - | 2 |
25 | Pterostichus (Petrophilus) sp. 1 | - | - | 2 | - | - | 2 |
- | Pterostichus spp. | - | - | 2 | 3 | - | 4 |
26 | Amara (Amara) sp. | - | - | - | 1 | - | 1 |
27 | Amara (Bradytus) aurichalcea Germar, 1823 | North | - | 1 | - | - | 1 |
28 | Curtonotus sp. | - | - | 4 | 3 | - | 4 |
29 | Agonum sp. | - | - | - | 1 | - | 1 |
30 | Olisthopus sturmii (Duftschmid, 1812) | + | - | - | 1 | - | 1 |
31 | Harpalus spp. | - | 2 | 3 | - | - | 3 |
32 | Cymindis cf. arctica Kryzhanovskij and Emetz, 1979 | East | 1 | 5 | - | - | 5 |
- | Cymindis spp. | - | 1 | - | 1 | - | 1 |
- | Carabidae indet. | - | 25 | - | 7 | 55 | - |
Hydrophilidae | |||||||
33 | Cymbiodyta marginella (Fabricius, 1792) | + | - | - | - | 1 | 1 |
34 | Cercyon sp. | - | - | - | 1 | - | 1 |
Helophoridae | |||||||
35 | Helophorus spp. | - | - | - | 6 | - | 3 |
Leiodidae | |||||||
36 | Leiodidae indet. | - | - | - | 1 | - | 1 |
Silphidae | |||||||
37 | Thanatophilus spp. | - | - | 1 | 4 | - | 2 |
- | Silphidae indet. | - | 1 | 5 | 1 | - | 5 |
Staphylinidae | |||||||
38 | Tachinus sp. | - | - | - | - | 1 | 1 |
39 | Omaliinae indet. | - | - | 1 | - | - | 1 |
- | Staphylinidae indet. | - | - | 3 | 1 | - | 3 |
Scarabaeidae | |||||||
40 | Aegialia cf. abdita Nikritin, 1975 | North | - | - | 1 | - | 1 |
41 | Aphodius multiplex Reitter, 1867 | South | 1 | - | - | - | 1 |
42 | Aphodius dictinctus (O.F. Müller, 1776) | + | - | - | 1 | - | 1 |
43 | Aphodius sp. | - | 4 | 7 | 15 | 2 | 8 |
Byrrhidae | |||||||
44 | Lamprobyrrhulus nitidus (Schaller, 1783) | + | 1 | 3 | 5 | 3 | |
45 | Morychus ostasiaticus Tshernyshev, 1997 | East | - | - | 4 | 2 | 2 |
46 | Porcinolus murinus (Fabricius, 1794) | South | - | - | 1 | - | 1 |
47 | Simplocaria elongata J. Sahlberg, 1903 | North | - | - | 1 | - | 1 |
48 | Cytilus sericeus (Foerster, 1771) | + | - | - | 1 | 1 | 1 |
Elateridae | |||||||
49 | Hypnoidus sp. | - | - | 1 | 3 | - | 2 |
50 | Elateridae indet. | - | - | - | 4 | 4 | 2 |
Coccinellidae | |||||||
51 | Coccinella sp. | - | - | 1 | - | 1 | 1 |
52 | Scymnus sp. | - | - | 1 | - | 1 | |
Tenebrionidae | |||||||
53 | Centorus rufipes (Gebler, 1833) | South | - | 7 | - | - | 7 |
Anobiidae | |||||||
54 | Xyletinus sp. | - | - | - | 1 | - | 1 |
Chrysomelidae | |||||||
55 | Hydrothassa hannoveriana (Fabricius, 1775) | North | - | - | 1 | - | 1 |
56 | Chrysomelidae indet. | - | - | 3 | 2 | 1 | 3 |
Brentidae | |||||||
57 | Mesotrichapion punctirostre (Gyllenhal in Schoenherr, 1839) | South | - | - | 3 | - | 2 |
58 | Apioninae indet. | - | - | - | 19 | 6 | 10 |
Curculionidae | |||||||
59 | Tournotaris bimaculata (Fabricius, 1787) | + | 13 | 19 | 12 | 23 | 19 |
60 | Notaris aethiops (Paykull, 1792) | North | 18 | 1 | - | 1 | 18 |
61 | Thryogenes nereis (Paykull, 1800) | + | 1 | 1 | 1 | - | 1 |
62 | Hylobius excavatus (Laicharting, 1781) | North | - | - | 1 | - | 1 |
63 | Pissodes insignatus Boheman in Schoenherr, 1843 | East | - | - | 1 | - | 1 |
64 | Lepyrus nordenskioldi Faust in Sahlberg, 1887 | North | 1 | - | - | - | 1 |
65 | Lepyrus sp. | - | - | - | - | 1 | 1 |
66 | Lixus paraplecticus (Linnaeus, 1758) | + | - | - | 2 | - | 1 |
67 | Stephanocleonus eruditus Faust, 1890 | East | 4 | - | 2 | - | 4 |
68 | Stephanocleonus favens Faust, 1884 | East | 1 | - | - | - | 1 |
69 | Stephanocleonus fossulatus (Fischer von Waldheim, 1823) | East | 1 | - | - | - | 1 |
70 | Stephanocleonus isochromus Suvorov, 1912 | East | 2 | - | - | - | 2 |
71 | Stephanocleonus leucopterus (Fischer von Waldheim, 1823) | South | 1 | - | - | - | 1 |
72 | Coniocleonus sp. | - | - | - | - | 1 | 1 |
- | Cleonini gen. sp. | - | - | - | 2 | - | - |
73 | Ceutorhynchus ignitus Germar, 1823 | + | - | - | 1 | - | 1 |
74 | Tychius albolineatus Motschulsky, 1860 | South | - | - | 1 | - | 1 |
75 | Hypera ornata (Capiomont, 1868) | North | - | 1 | - | - | 1 |
Hypera sp. | - | - | - | - | 1 | 1 | |
76 | Metadonus distinguendus (Boheman in Schoenherr, 1842) | + | 1 | 1 | - | - | 1 |
77 | Trichalophus biguttatus (Gebler, 1832) | East | 1 | - | - | - | 1 |
78 | Sitona sp. | - | 2 | 14 | - | - | 14 |
79 | Chlorophanus sibiricus Gyllenhal in Schoenherr, 1834 | + | - | - | - | 1 | 1 |
80 | Phyllobius femoralis Boheman in Schoenherr, 1842 | East | 1 | - | - | - | 1 |
81 | Phyllobius virideaeris (Laicharting, 1781) | + | 3 | 1 | 2 | - | 3 |
Phyllobius spp. | - | - | 4 | 14 | 6 | 7 | |
82 | Polydrusus amoenus (Germar, 1823) | North | 7 | - | - | - | 7 |
83 | Otiorhynchus arcticus (O. Fabricius, 1780) | North | - | 1 | - | - | 1 |
84 | Otiorhynchus beatus Faust, 1890 | East | - | - | 2 | - | 1 |
85 | Otiorhynchus grandineus Germar, 1823 | East | - | - | 2 | - | 1 |
86 | Otiorhynchus janovskii Korotyaev, 1990 | East | - | 2 | 13 | - | 7 |
87 | Otiorhynchus altaicus Stierlin, 1861 | South | 220 | 165 | 103 | 82 | 220 |
88 | Otiorhynchus ursus Gebler, 1844 | South | 13 | ||||
89 | Otiorhynchus obscurus Gyllenhal in Schoenherr, 1834 | South | 19 | 41 | 13 | 11 | 41 |
90 | Otiorhynchus politus Gyllenhal in Schoenherr, 1834 | North | 4 | 11 | 6 | 18 | 11 |
91 | Otiorhynchus pullus Gyllenhal in Schoenherr, 1834 | South | 1 | 6 | 2 | - | 6 |
92 | Otiorhynchus unctuosus Germar, 1823 | South | - | 1 | 11 | - | 6 |
- | Otiorhynchus sp. | - | - | - | - | 19 | - |
- | Coleoptera indet. | - | 17 | 3 | - | 5 | - |
- | Hymenoptera indet. | - | 5 | - | - | 7 | 5 |
* Modern distribution in comparison with the Dubrovino site.