TABLE 1. Summary of the morphological features, which are useful to distinguish between the species of Eosemionotus.
Morphological features |
E. vogeli |
E. ceresiensis |
E. diskosomus |
E. sceltrichensis |
E. minutus |
Body shape | fusiform | fusiform | deep | deep | fusiform |
Ornamented dermal skull bones | present | absent | absent | absent | absent |
Length frontal process | ? | short | long | long | long |
Relationship between frontal processes | ? | coalescent | coalescent | coalescent | separate |
Supraoccipital canal in parietals | present | present | absent | absent | present |
Parietals fused | absent | absent | absent | present | absent |
Number of extrascapulars | 1 pair | 5 | 3 pairs | 3 pairs | 3 pairs |
Maximal number of premaxillary teeth | 4 | 6 | 3-4 | 4 | 6 |
Number of dentary teeth | 7 | 10-12 | ? | ? | 11 |
Maxillary teeth | ? | present | absent | absent | present? |
Shape of maxilla | slender | slender | deep | deep | slender |
Position of lower jaw articulation relative to orbit | centre | anterior | anterior | centre | centre |
Depth to length ratio of opercle | ~1 | ~1 | >1 | >1 | >1 |
Length of interopercle | short | short | short | long | ? |
Number of branchiostegals | 6-7 | 7-8 | 5 | 6 | 5 |
Number of caudal fin rays | 12-15 | 13-15 | 13 | 12 | 12 |
Number of caudal fin rays below the lateral line | 7-8 | 8-9 | 7 | 8 | 8 |
Size of fringing fulcra | slender | small | large | large | slender |
Fringing fulcra on dorsal margin of caudal fin | present | present | present | present | absent |
Number of scales in marginal row of body lobe | 6-7 | 5 | 4 | 2 | 4 |
Number of scales posterodorsal to the hinge-line | 4 | 5 | 4 | 4-5 | 3 |
Number of dorsal caudal fulcra on the body lobe | 5-6 | 5 | 4 | 4 | 4 |
Number of vertical rows of scales | 26-28 | 27-28 | 23-25 | 23-25 | 24 |
Dorsal precaudal scutes | ? | 10 | 7-8 | absent | 1 |
Ventral precaudal scutes | absent | absent | absent | 4 | 3 |
Number of anal fin rays | 7 | 6 | 6-7 | 7 | 5 |
Number of dorsal fin rays | 6-10 | 6 | 9 | 6-7 | 6 |
TABLE 2. Body proportions based on measurements taken on well-preserved specimens. SL: standard length, HL: head length, BD: maximal body depth, CD: minimal depth of the caudal peduncle.
Species | Specimen | SL | HL/SL | BD/SL | CD/SL |
E. ceresiensis | PIMUZ T357 | 39,00 | 0,35 | 0,33 | 0,13 |
PIMUZ T236 | 38,00 | 0,30 | 0,41 | 0,15 | |
PIMUZ T319 | 42,50 | 0,32 | 0,42 | 0,18 | |
PIMUZ T329 | 36,50 | 0,33 | 0,42 | 0,18 | |
E. diskosomus | MCSN 8082 | 45,00 | 0,37 | 0,58 | 0,20 |
MCSN 8006 | 60,50 | 0,35 | 0,51 | 0,17 | |
MCSN 5617 | 46,70 | 0,36 | 0,61 | 0,20 | |
PIMUZ T2924 | 22,50 | 0,39 | 0,55 | 0,20 | |
E. sceltrichensis | MCSN 8418 | 52,80 | 0,48 | ||
MCSN 8497 | 39,14 | 0,45 | |||
MCSN 8491 | 33,16 | 0,53 | |||
E. minutus | MCSN 8482 | 25,00 | 0,39 | 0,38 | 0,18 |
TABLE 3. Fossil record of Eosemionotus.
Taxonomy | Stratigraphic provenance | Geographic provenance | Reference |
Eosemionotus sp. | Carnian, Pyrenean basin | Vilanova de la Sal, Lleida, Spain | Fortuny et al., 2011 |
Eosemionotus sp. | upper Ladinian, Falang Fm. (Zhuganpo Member) | Changdi, Luoping, Yunnan, China | Jin, 2006 |
Eosemionotus sp. | upper Ladinian, Alcover unit | Alcover, Tarragona, Spain | Cartanyà, 1999 |
E. ceresiensis, E. diskosomus, E. sceltrichensis, E. minutus |
Ladinian, Meride Limestone | Several localities, Monte San Giorgio, Switzerland | this work |
Eosemionotus sp. | lower Ladinian, Prosanto Fm. | Ducan and Landwasser areas, Graubünden, Switzerland | Bürgin et al., 1991; Herzog, 2003 |
E. ceresiensis, E. diskosomus |
lowermost Ladinian, Besano Fm. | Several localities, Monte San Giorgio, Switzerland | Bürgin, 2004; this work |
Eosemionotus sp. | upper Anisian, Guanling Fm. (Member II) | Xinmin, Panxian, Guizhou, China | Jin, 2006 |
E. vogeli | upper Anisian, Diemel Fm. (Middle Muschelkalk) | Grebenberg bei Angerstein, Germany | Schultze and Möller, 1986 |
E. vogeli | upper Anisian, Heilbronn Formation, ‘blaue Fischmergel’ of Rüdersdorf (Middle Muschelkalk) | Rüdersdorf bei Berlin, Germany | Siegel and Wischnewsky, 2017; this work |
E. vogeli | upper Anisian, Karlstadt Fm. (Middle Muschelkalk) | Förderstedt bei Staßfurt, Germany | Fritsch, 1906 |
Eosemionotus sp. | upper Anisian, Strelovec Fm. | Robanov kot valley, Kamnik-Savinja Alps, Slovenia | Miklavc et al., 2016 |
Eosemionotus sp. | middle Anisian, Velika Planina Horizon | Kamnik-Savinja Alps, Slovenia | Hitij et al., 2010 |
Eosemionotus sp. | middle Anisian, Upper Member, Guanling Fm. | Dawazi site, Yunnan, China | Sun et al., 2009 |
Eosemionotus sp. | middle - upper Anisian, Vossenveld Fm. (Lower Muschelkalk) | Steen- en Kalkgroeve Winterswijk, The Netherlands | Oosterink and Poppe, 1979; Hagdorn and Simon, 2010; Maxwell et al., 2016 |