TABLE 1. Age, tectonic and depositional settings of studied geological formations. Modified from Mihaljević et al., 2017.
Geological Formation | Age | Tectonic setting | Depositional setting | References |
Butong Limestone Formation (Cebu) | late Oligocene - early Miocene | Backarc setting? Oblique convergence | Isolated low relief buildups | Porth et al., 1989; Wilson, 2002; Aurelio and Peña, 2010. |
Calagasan Formation (Cebu) |
late Oligocene | Overlying the Butong Limestone | Coral rich limestone lenses within a shale | Aurelio and Peña, 2010; Mihaljević et al., 2017 |
Melinau Limestone (Sarawak) |
Eocene - early Miocene | Isolated carbonate platform, basin margin | Carbonate platform, some marginal deposits, coral patch reefs only noted in Miocene | Adams and Haak, 1962; Adams, 1965; Wannier, 2009; Mihaljević et al., 2014 |
Subis Limestone (Sarawak) |
early Miocene | Baram Balabac Basin margin | Isolated carbonate platform formed on palaeohigh | Adams, 1965; Mihaljević et al., 2014 |
Trankalan/Binaguiohan Limestone (Negros) | late Oligocene - early Miocene | Unclear | Carbonate platform with localized patch reefs | Jurgan and Domingo, 1989; Porth et al., 1989; Aurelio and Peña, 2010 |
TABLE 2. An overview of the fossil collection localities with geological formation, their age and latitude and longitude coordinates. From Mihaljević et al., 2017.
Locality | Geological Formation | Age | Lat/long |
CF1 | Calagasan Formation | late Oligocene (middle Chattian) |
N 9°54'22.9'' E 123°29'56'' |
CF2 | Calagasan Formation | late Oligocene (middle Chattian) |
N 9°53'42.3'' E 123°29'46.2'' |
CF4 | Calagasan Formation | late Oligocene (middle Chattian) |
N 9°53'0.49'' E 123°29'49.33'' |
CF5 | Calagasan Formation | late Oligocene (middle Chattian) |
N 9°53'13.3'' E 123°29'49.5'' |
CF6 | Calagasan Formation | late Oligocene (middle Chattian) |
N 9°54'7.4'' E 123°29'44.8'' |
CF7 | Calagasan Formation | late Oligocene (middle Chattian) |
N 9°54'7.1'' E 123°30'5.5'' |
CF10 | Calagasan Formation | late Oligocene (middle Chattian) |
N 9°53'13'' E 123° 29' 49.75'' |
BLF1 | Butong Limestone Formation | late Oligocene (late Chattian) |
N 9°54'23.2'' E 123° 29' 51.5'' |
BLF2 | Butong Limestone Formation | late Oligocene (late Chattian) |
N 9°54'20.6'' E 123°29'53.8'' |
BLF4 | Butong Limestone Formation | late Oligocene (late Chattian) |
N 9°54'37.7'' E 123°30'4'' |
TBL | Trankalan/Binaguiohan Limestone | early Miocene (Aquitanian) |
N 9°48'7.89'' E 122°37'17.56'' |
ML | Melinau Limestone | early Miocene (Aquitanian) |
N 4° 1' 57.9" E 114° 47' 19° |
SL1 | Subis Limestone | early Miocene (Aquitanian) |
N 3° 46' 45.66" E 113° 46' 50.95" |
SL2 | Subis Limestone | early Miocene (Aquitanian) |
N 3° 46' 42.82" E 113° 47' 15.68" |
TABLE 3. Comparison of taxonomic differences between morphospecies within genus Astrea.
Morphospecies | Colony shape | Average corallite width (mm) |
Coenosteum amount | Number of septa per 5 mm |
Preservation remarks |
Astrea sp. A | encrusting to massive | 7.72 - 11.37 | moderate (<corallite diameter) | eight to 11 | no relief preserved |
Astrea sp. B | encrusting | 4.47 - 9.28 | moderate (<corallite diameter) but more widely than Astrea sp. 1 | eight to 11 | slightly (< 1 mm) elevated walls |
Astrea sp. C | massive | 5.19 - 10.73 | moderate (<corallite diameter) | six to 11 | some relief preserved, poorly |
TABLE 4. Comparison of taxonomic differences between morphospecies within genus Dipsastraea.
Morphospecies | Colony shape |
Average corallite width (mm) |
Corallite outline | Corallite relief | Paliform lobes |
Dipsastraea cf. speciosa | massive | 4.22 - 9.3 | round | ~3 | weak to moderate |
Dipsastraea sp. A | massive | 4.67 - 6.34 | tubular, exsert | 3.05 - 5.65 | weak to moderate |
Dipsastraea sp. B | encrusting to massive | 5.94 - 13.64 | irregular | 3.23 - 5.51 | weak |
TABLE 5. Comparison of taxonomic differences between morphospecies within genus Favites.
Morphospecies | Colony shape | Average corallite width (mm) |
Corallite discrete | Corallite outline | Number of septa per 5 mm |
Favites cf. chinensis | encrusting to massive | 10.94 - 14.16 | monocentric | polygonal and irregular | seven to 11 |
Favites cf. pentagona tenuis | encrusting to massive | 6.07 - 10.35 | one to three centers | irregular | six to ?24 |
Favites sp. A | massive | 6.70 - 9.26 | one to three centers | round to elliptical | six to 11 |
Favites sp. B | massive | 4.14 - 9.12 | monocentric | rounded and regular | eight to 11 |
TABLE 6. Comparison of taxonomic differences between morphospecies within genus Goniopora.
Morphospecies | Colony shape | Branch diameter | Average corallite width (mm) |
Corallite outline | Paliform lobes |
Goniopora cf. palmensis | branching | 8.55 - 19.67 | 2.21 - 3.66 | mostly round | prominent |
Goniopora cf. planulata | branching to columnar | 14.09 - 23.88 | 2.73 - 4.14 | mostly round or irregular | moderate |
Goniopora cf. tenuidens | massive | NA | 3.87 - 7.80 | mostly round | weak to moderate |
TABLE 7. Checklist of described material from Malaysia and the Philippines and the formation in which they were found. Numbers represent number of museum lots. Geologic formation abbreviations: CF - Calagasan Formation, BLF - Butong Limestone Formation, TBL - Trankalan/Binaguiohan Limestone, ML - Melinau Limestone, SL - Subis Limestone.
Taxon | CF | BLF | TBL | ML | SL |
Acanthastrea cf. echinata | 3 | 1 | 0 | 1 | 0 |
Acanthastrea sp. A | 2 | 0 | 0 | 0 | 0 |
Acropora sp. A | 1 | 7 | 0 | 0 | 0 |
Acropora sp. B | 5 | 5 | 0 | 0 | 0 |
Actinastrea cf. minutissima | 38 | 24 | 0 | 0 | 0 |
Alveopora cf. fenestrata | 0 | 0 | 0 | 2 | 0 |
Astrea sp. A | 2 | 0 | 0 | 0 | 0 |
Astrea sp. B | 0 | 1 | 0 | 0 | 0 |
Astrea sp. C | 0 | 0 | 0 | 0 | 2 |
Astreopora cf. digitata | 0 | 0 | 0 | 0 | 9 |
Astreopora cf. expansa | 0 | 1 | 0 | 0 | 0 |
Blastomussa cf. simplicitexta | 0 | 0 | 1 | 0 | 0 |
Coelastrea sp. A | 0 | 4 | 0 | 0 | 0 |
Cyphastrea cf. serailia | 4 | 9 | 0 | 0 | 0 |
Cyphastrea sp. A | 1 | 0 | 0 | 0 | 0 |
Dictyaraea cf. anomala | 0 | 1 | 0 | 0 | 0 |
Dictyaraea cf. micrantha | 1 | 4 | 0 | 0 | 0 |
Dipsastraea cf. speciosa | 3 | 2 | 2 | 0 | 23 |
Dipsastraea sp. A | 4 | 0 | 0 | 0 | 0 |
Dipsastraea sp. B | 0 | 1 | 0 | 0 | 0 |
Favites cf. chinensis | 6 | 1 | 0 | 0 | 0 |
Favites cf. pentagona tenuis | 1 | 0 | 0 | 0 | 0 |
Favites sp. A | 5 | 0 | 0 | 0 | 0 |
Favites sp. B | 2 | 1 | 0 | 0 | 0 |
Favites spp. | 0 | 0 | 0 | 7 | 14 |
Fungia spp. | 3 | 0 | 0 | 1 | 0 |
Galaxea sp. A | 3 | 0 | 0 | 0 | 2 |
Gardineroseris cf. planulata | 0 | 1 | 0 | 0 | 0 |
Goniastrea cf. stelligera | 1 | 9 | 0 | 0 | 0 |
Goniopora cf. palmensis | 2 | 7 | 0 | 0 | 0 |
Goniopora cf. planulata | 5 | 8 | 0 | 0 | 0 |
Goniopora cf. tenuidens | 1 | 0 | 0 | 0 | 0 |
Hydnophora cf. microconos | 11 | 1 | 0 | 15 | 14 |
Leptastrea cf. transversa | 11 | 1 | 0 | 0 | 0 |
Leptastrea sp. A | 4 | 0 | 0 | 0 | 0 |
Leptoria cf. phrygia | 0 | 1 | 0 | 7 | 2 |
Leptoseris sp. A | 0 | 1 | 0 | 0 | 0 |
Lobophyllia cf. hemprichii | 4 | 0 | 0 | 0 | 0 |
Pachyseris spp. | 1 | 58 | 0 | 0 | 0 |
Paramontastraea sp. A | 0 | 0 | 0 | 3 | 0 |
Pavona cf. clavus | 1 | 0 | 0 | 0 | 2 |
Pavona cf. venosa | 0 | 7 | 0 | 0 | 0 |
Platygyra cf. lamellina | 25 | 1 | 2 | 0 | 1 |
Porites sp. A | 0 | 0 | 1 | 0 | 0 |
Porites spp. | 1 | 4 | 2 | 0 | 6 |
Stylophora cf. pistillata | 4 | 13 | 0 | 0 | 0 |
Stylophora spp. | 1 | 7 | 0 | 1 | 0 |
Trachyphyllia cf. constricta | 5 | 0 | 0 | 0 | 0 |
Trochocyathus cf. schmidti | 1 | 1 | 0 | 0 | 0 |