TABLE 1. Taxonomic study history of palaeolenid trilobites from the Guanshan Biota.
Publication | Species considered | Relevant notes |
Mansuy (1912, p. 27) | Palaeolenus douvillei, P. deprati and P. “lantenoisi” | First description of three Palaeolenus species using a few specimens (n=6) with all the cranidial features exhibiting |
Chang (1966, p. 150). | Megapalaeolenus deprati | Moved Palaeolenus deprati into Megapalaeolenus |
Chang et al. (1980, p. 234, pl. 73, Figs 9-12) | Palaeolenus longispinus | Erected Palaeolenus longispinus using robust pleural spines on second thoracic tergite |
Lin and Peng (2004) | Megapalaeolenus deprati and Palaeolenus douvillei | Doubted the validity of Megapalaeolenus. Noted that holotype was deformed and considered M. deprati conspecific with Palaeolenus douvillei |
Luo et al. (2007) | Megapalaeolenus deprati, Palaeolenus douvillei and P. “lantenoisi” | Considered all species and genera to be valid |
Lin and Peng (2009) | Megapalaeolenus deprati and Palaeolenus douvillei | Considered species conspecific |
Hu et al. (2013) | Megapalaeolenus deprati, Palaeolenus douvillei, P. “lantenoisi” and P. longispinus | Suggested all taxa were valid species |
Lin (2014) | All the published 34 specimens assigned to palaeolenids (n=34) | Used cluster analysis to group all taxa into Palaeolenus. Suggested P. douvillei and Megapalaeolenus deprati conspecific |
Luo et al. (2014) | Palaeolenus douvillei and Megapalaeolenus deprati | Considered both taxa valid. Noted they are found in different members of the Wulongqing Formation |
Lin and Peng (2016) | All the published 34 specimens assigned to palaeolenids (n=34) | Used cluster analysis to group. Reconfirmed Lin (2014), suggesting Palaeolenus douvillei and Megapalaeolenus deprati are conspecific |
TABLE 2. Locations of cephalic landmarks for analyzed specimens.
Landmark | Description |
1 | Intersection of anterior border with sagittal line |
2 | Intersection of border furrow with sagittal line |
3 | Intersection of anterior margin of frontal lobe with sagittal line |
4 | Intersection of anterior margin of occipital ring with sagittal line |
5 | Intersection of posterior margin of occipital ring with sagittal line |
6 | Intersection of axial glabellar furrow and posterior margin |
7 | Anterior border of palpebral lobe |
8 | Posterior border of palpebral lobe |