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TABLE 1. Comparison of critical morphological characteristics of fossils of the genus Dilophus.

  D. idanos D. crassicornis D. palaeofebrilis D. succineus D. pumilio D. matilei D. krantzii D. luteipennis D. andrewrossi D. pinguis
Locality Montana Baltic Baltic Baltic Germany Dominican Republic Germany France England Croatia
Age Middle Eocene Late
Eocene
Late
Eocene
Late
Eocene
Miocene Miocene Oligocene Oligocene Late Eocene Miocene
Matrix Shale Amber Amber Amber Shale Amber Shale Shale Limestone Limestone
Sex M M & F M M & F F M M F   M & F
Body length (mm) 3.7 3.6-3.7 (M);
4.3-5.1 (F)
4.3-5.7 3.6 (M);
3.5 (F)
4.5 3.8 11 5.5-6.0 ND 12.6 (M);
15 (F)
Head length (mm) 0.52 0.57-0.62 (M);
0.62-0.77 (F)
0.78-0.92 0.92 (M);
0.62 (F)
0.8 0.65 ND 0.7 ND ND (M);
2.0 (F)
Palpus length (mm) ND ND 0.38 ND ND 0.56 twice as long as flagellum ND ND ND
Antenna length (mm) 0.2 0.15
(M; flagellum)
0.5 0.48 (M);
0.50
(F)
ND 0.53 ND ND ND ND
Number of flagellomeres 6 6-7 (M); 7 (F) 12 12
(M & F)
6? 7 at least 8 ND ND ND (M);
“possibly six” (F)
Thorax L/W (mm) 0.93/0.60 0.92-0.99/0.62 (M);
1.22-1.55/? (F)
1.31-1.52/1.07-1.15 1.2/0.64
(F)
1.4/? 1.1/0.59 ND 1.3-1.5/1.4 ND 3.2 (M);
5.6 (F)
Wing L/W (mm) 2.4/0.91 3.2-3.7/1.2
(M):
3.75/? (F)
3.55/1.10 3.8/1.39
(F)
3.8/1.5 2/0.8 L = ̴8.6 mm (Fig.) or "about length of body" (text) 3.8-4.5 (4.0-4.6)/1.5-1.7 (1.5-1.8) 3.85/1.45 9.5/3.3 (M); 12.3/4.4 (F)
r-m:base of Rs 3.2 2.1-2.5 (M);
1.3 (F)
3 ND 2.1 4.5 0.47 0.6? 2 ND (M); 1 (F)
Fore tibial spines 2:2:8 3:6 (M);
3:6 (F)
1:2:8 2:3:6
(M & F)
2:2:? or 2:3:? 2:4:8 Apex with ring of spines 3? mesal ?:?:2 2? mesally, small; fine & short apically (M); 3? mesally; 6? in circlet, short (F)
Mesonotal spines 12 (anterior)
and 10
(posterior) =
12 + 10
10 + ? (M);
8 + 2 + 8 (F)
10-11 + 10 ? + 10 (M);
10 + 10 (F)
Vague
trace
2 rows of
10-12 each
ND Not visible ND ND (M); “short, stout (F)
Abdomen L/W (mm) 2.3/0.45 ND ND ND 3.0/? 2.14/0.52 6/? 3.7-4.4/1.5 ND 8.3/2.1 (M); 10.1/ND (F)
Reference Greenwalt and
Fitzgerald, 2022
Skartveit, 2008 Skartveit, 2008 Skartveit, 2008 Skartveit and Pika, 2014 Waller, Nel and Menier, 2000 Heyden, 1870 Théobald, 1937;
Skartveit and Nel, 2017
Krzemiński et al., 2019 Heer, 1849;
Skartveit & Krizmanic, 2020