APPENDIX 1
Figure illustrating the locations of measurements of Uncinidae chelae morphology used in this study.
APPENDIX 2
Table of measurements of Uncinidae chelae morphology.
Specimen Number | Species | Figure in Appendix 4 |
Manus Length (mm) |
Manus Height (mm) |
Propodus Length (mm) |
Dactylus Length (mm) |
Dactylus Thickness (mm) |
Dactylus Height (mm) |
Manus Height at Attachment (mm) |
Index Length (mm) |
Notch Length (mm) |
Notch Height (mm) |
Index Height (mm) |
Index Length to Index Height Ratio |
Notch Length to Notch Height Ratio |
Notch Height to Notch Length Ratio |
Manus Length to Index Length Ratio |
Manus Height at Attachment to Manus Height Ratio |
Dactylus Thickness to Index Height Ratio |
TMP 2018.024.0005 | U. pacifica | A4-26 | - | - | - | 23.01 | 11.28 | 16.92 | - | - | - | - | - | - | - | - | - | - | - |
TMP 2018.024.0035 | U. pacifica | A4-31 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - |
TMP 2018.024.0004 | U. pacifica | A4-34 | - | 10.60 | - | - | - | - | - | 7.83 | 2.81 | 3.88 | 6.18 | 1.267 | 0.724 | 1.381 | - | - | - |
TMP 2018.024.0003 | U. pacifica | A4-27 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - |
TMP 2017.015.0007 | U. pacifica | A4-25 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - |
TMP 2017.015.0005 | U. pacifica | A4-24 | 27.04 | 11.85 | 38.33 | 12.61 | 4.64 | 8.61 | 5.61 | 11.29 | 3.2 | 3.28 | 7.32 | 1.542 | 0.976 | 1.025 | 2.395 | 0.473 | 0.634 |
TMP 2017.015.0003 | U. pacifica | A4-23 | 14.17 | 8.99 | 25.46 | 11.15 | 4.28 | 7.46 | - | - | - | - | - | - | - | - | - | - | - |
TMP 2017.015.0001a | U. pacifica | A4-22 | 16.92 | 9.59 | 24.5 | 9.81 | 4.01 | 7.26 | 4.76 | 6.46 | 2.39 | 2.91 | 6.69 | 0.966 | 0.821 | 1.218 | 2.619 | 0.496 | 0.599 |
TMP 2017.015.0001b | U. pacifica | A4-22 | 15.21 | 10.22 | 24.69 | 10.42 | 4.53 | 6.47 | 5.02 | 5.05 | 2.42 | 3.65 | 7.38 | 0.684 | 0.663 | 1.508 | 3.012 | 0.491 | 0.614 |
TMP 2016.027.0003 | U. pacifica | A4-21 | 34.87 | 22.91 | - | - | - | - | 9.81 | - | - | - | 16.53 | - | - | - | - | 0.428 | - |
TMP 2015.050.0069 | U. pacifica | A4-18 & A4- 19 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - |
TMP 2015.050.0039 | U. pacifica | A4-17 | 27.63 | 17.38 | 15.7 | 7.36 | 9.45 | 7.36 | 11.6 | - | - | 9.6 | 1.208 | - | - | 2.382 | 0.423 | 0.767 | |
TMP 2014.021.0023 | U. pacifica | A4-39 | 26.82 | 19.62 | 45.43 | 21.83 | 9.99 | 13.89 | 7.92 | 14.84 | 3.39 | 6.74 | 14.94 | 0.993 | 0.503 | 1.988 | 1.807 | 0.404 | 0.669 |
TMP 2015.050.0038 | U. pacifica | A4-16 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - |
TMP 2014.021.0033 | U. pacifica | A4-13 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | |
TMP 2014.021.0034 | U. pacifica | A4-14 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - |
TMP 2014.021.0007 | U. pacifica | A4-12 | 31.05 | 15.67 | 49.76 | 15.75 | 6.95 | 7.47 | 5.74 | 10.79 | - | 4.91 | 12.16 | 0.887 | - | - | 2.878 | 0.366 | 0.572 |
TMP 2002.043.0012 | U. pacifica | A4-7 | 12.11 | 5.56 | 7.74 | - | - | - | - | - | - | - | - | - | - | - | |||
TMP 2002 043.0019a | U. pacifica | A4-9 | 14.91 | 8.28 | 27.7 | 11.2 | 4.47 | 8.04 | 4.98 | 7.02 | 2.78 | 2.71 | 6.37 | 1.102 | 1.026 | 0.975 | 2.124 | 0.601 | 0.702 |
TMP 2002 043.0019b | U. pacifica | A4-9 | - | - | - | 11.8 | 4.24 | 8.67 | - | - | - | - | - | - | - | - | - | - | - |
TMP 2014.021.0003 | U. pacifica | A4-10 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - |
TMP 2002.043.0008 | U. pacifica | A4-6 | 18.02 | 9.5 | 27.7 | - | - | - | 4.35 | 8.77 | 3.26 | 2.84 | 6.98 | 1.256 | 1.148 | 0.871 | 2.055 | 0.458 | - |
TMP 2002.043.0015 | U. pacifica | A4-8 | - | - | - | - | - | - | - | 7.4 | - | - | 6.97 | 1.062 | - | - | - | - | - |
TMP 2002.043.0007 | U. pacifica | A4-5 | - | - | - | 10.75 | 4.48 | 7.8 | - | - | - | - | - | - | - | - | - | - | - |
TMP 2002.043.0003a | U. pacifica | A4-2 | - | 9.63 | - | - | - | - | - | 8.38 | 4.07 | 4.1 | 8.59 | 0.976 | 0.993 | 1.007 | - | - | - |
TMP 2002.043.0003b | U. pacifica | A4-2 | - | - | - | - | - | - | - | 7.96 | 2.69 | 4.12 | 7.19 | 1.107 | 0.653 | 1.532 | - | - | - |
TMP 2002.043.0006 | U. pacifica | A4-4 | - | 8.79 | - | - | - | - | - | 6.35 | 2.68 | 3.25 | - | - | 0.825 | 1.213 | - | - | - |
TMP 2002.043.0002 | U. pacifica | A4-1 | - | - | - | 16.56 | 6.9 | 10.78 | - | - | - | - | - | - | - | - | - | - | - |
TMP 2014.021.0008 | U. pacifica | A4-38 | - | - | - | 15.33 | 7.32 | 11.51 | - | - | - | - | - | - | - | - | - | - | - |
TMP 2014.021.0002 & TMP 2013.036.0004 | U. pacifica | A4-37 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - |
TMP 2002.043.0004 | U. pacifica | A4-3 | 16.63 | 11.34 | 27.68 | 13.15 | 4.76 | 7.21 | 4.78 | 8.69 | 3.55 | 3.35 | 6.9 | 1.259 | 1.060 | 0.944 | 1.914 | 0.422 | 0.690 |
TMP 2002.043.0005 | U. pacifica | A4-35 | - | 19.94 | - | 18.22 | 8.32 | 12.68 | - | 10.73 | 4.34 | 6.44 | - | - | 0.674 | 1.484 | - | - | - |
TMP 2014.021.0004 | U. pacifica | A4-11 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - |
KMNH IvP 300,027 | U. pacifica | Schweigert et al., 2003, Figure 4 | 26.43 | - | 29.98 | 11.08 | 28.073 | - | 20.89 | 4.4 | 4.54 | 7.9 | - | - | 0.575 | 1.740 | 6.007 | - | - |
TMP 2018.024.0009 | U. ollerenshawi | A4-30 | - | - | - | 25.6 | 7.38 | 10.75 | - | - | - | - | - | - | - | - | - | - | |
TMP 2018.024.0006 | U. ollerenshawi | A4-28 | - | - | - | - | - | - | - | - | 4.75 | 3.53 | 9.07 | - | 1.346 | 0.743 | - | - | - |
TMP 2016.027.0008 | U. ollerenshawi | A4-33 | - | - | - | 41.54 | 8.95 | 15.99 | - | - | - | - | - | - | - | - | - | - | |
TMP 2015.051.0063 | U. ollerenshawi | A4-20 | 19.54 | 10.71 | 41.46 | 21.04 | 5.9 | 8.24 | 7.54 | 17.84 | 2.03 | 3.14 | 7.05 | 0.395 | 0.646 | 0.646 | 1.095 | 0.704 | 0.837 |
TMP 2014.021.0052A | U. ollerenshawi | A4-15 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | |
TMP 2014.021.0052B | U. ollerenshawi | A4-15 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | |
TMP 2015.050.0011 | U. ollerenshawi | A4-40 | 24.77 | 11.5 | 53.09 | 29.16 | 5.84 | 8.86 | 9.06 | 21.4 | 6.65 | 7.22 | 7.02 | 0.328 | 0.921 | 0.921 | 1.157 | 0.788 | 0.832 |
TMP 2013.036.0002 | U. ollerenshawi | A4-16 | 35.2 | 17.13 | 74.76 | 37.94 | 8.7 | 13.04 | 10.63 | 31.66 | 5.35 | 4.69 | 11.18 | 0.353 | 1.141 | 1.141 | 1.112 | 0.621 | 0.778 |
TMP 2005.028.0004 | U. ollerenshawi | A4-42 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - |
TMP 2018.024.0007 | U. ollerenshawi | A4-29 | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - | - |
GSC 80067 A * HOLOTYPE | U. ollerenshawi | A4-32 | 43.67 | 12.05 | 64.61 | 22.11 | 5.84 | 11.31 | 8.81 | 13.95 | 4.84 | 4.959 | 8.33 | 0.597 | 0.976 | 0.976 | 3.130 | 0.731 | 0.701 |
GSC 80067 B * HOLOTYPE | U. ollerenshawi | A4-32 | 40.54 | 14.49 | 59.41 | 22.01 | 6.4 | 11.1 | 8.65 | 14.27 | 5.18 | 4.5 | 6.8 | 0.477 | 1.151 | 1.151 | 2.841 | 0.597 | 0.941 |
MB.A.3826 top claw | U. posidoniae | A4-41 | 24.73 | 7.67 | 39.16 | 14.77 | 4.08 | 7.06 | 5.03 | 9.9 | 2.43 | 3.06 | 4.07 | 0.411 | 0.794 | 1.259 | 2.498 | 0.656 | 1.002 |
MB.A.3826 bottom claw | U. posidoniae | A4-41 | 24.52 | 7.99 | 39.51 | 13.27 | 3.56 | 6.61 | 5.24 | 9.8 | 2.39 | 2.94 | 4.74 | 0.484 | 0.813 | 1.612 | 2.502 | 0.656 | 0.751 |
Posidoniae1A | U. posidoniae | 46.58 | 16.731 | 70.275 | 27.12 | 7.46 | 9.846 | 14.098 | 20.376 | - | - | 7.797 | 0.383 | - | - | 2.286 | 0.843 | 0.957 | |
Posidoniae1B | U. posidoniae | 44.228 | 17.3 | 70.89 | 28.049 | 14.77 | 9.45 | 14.6 | 19.608 | 5.61 | 4.98 | 9.57 | 0.488 | 1.127 | 1.922 | 2.256 | 0.844 | 1.543 | |
Posidoniae2 | U. posidoniae | 54.74 | 19.62 | 91.92 | 32.16 | 10.82 | 17.72 | 11.009 | 22.61 | 5.18 | 7.06 | 13.7 | 0.606 | 0.734 | 1.941 | 2.421 | 0.561 | 0.790 | |
Poidoniae LECTOTYPE | U. posidoniae | Schweigert et al., 2003, Plate 1 | 57.96 | 20.167 | 91.18 | 32.404 | 10.77 | 16.064 | 13.971 | 23.506 | - | - | 12.278 | 0.522 | - | - | 2.466 | 0.693 | 0.877 |
SMNS no. 29150 top claw | U. posidoniae | Schweigert et al., 2003, Plate 2 | 45.81 | 16.12 | 69.31 | 27.496 | 9.573 | 14.533 | 14.134 | 19.006 | - | - | 8.967 | 0.472 | - | - | 2.410 | 0.877 | 1.068 |
SMNS no. 29150 bottom claw | U. posidoniae | Schweigert et al., 2003, Plate 3 | 45.66 | 16.26 | 68.54 | 26.419 | 9.451 | 14.22 | 13.802 | 18.971 | - | - | 9.097 | 0.480 | - | - | 2.407 | 0.849 | 1.039 |
No ID "Plate 4" top claw | U. posidoniae | Schweigert et al., 2003, Plate 4 | 40.495 | 11.034 | 63.584 | 22.9 | 5.935 | 10.932 | 8.661 | 14.718 | 4.754 | 4.566 | 8.038 | 0.546 | 1.041 | 1.760 | 2.751 | 0.785 | 0.738 |
No ID "Plate 4" bottom claw | U. posidoniae | Schweigert et al., 2003, Plate 4 | 41.204 | 12.804 | 62.095 | 21.799 | 5.517 | 12.36 | 8.96 | 14.705 | 3.799 | 4.857 | 6.503 | 0.442 | 0.782 | 1.339 | 2.802 | 0.700 | 0.848 |
SMNS no. 62815 | U. posidoniae | Schweigert et al., 2003, Plate 8 | 45.135 | 14.623 | 67.417 | 23.006 | 6.774 | 11.284 | 11.065 | 14.493 | 4.549 | 5.562 | 9.162 | 0.632 | 0.818 | 1.647 | 3.114 | 0.757 | 0.739 |
MSNM no. i10864 left claw | U. alpina | Schweigert et al., 2003, Plate 10, Figure 2 | 13.31 | 3.15 | 19.08 | 5.12 | 2.03 | 2.74 | 2.84 | 1.65 | 0.79 | 1.72 | 1.99 | 1.20 | 0.46 | 2.18 | 8.06 | 0.90 | 1.02 |
MSNM no. i10864 right claw | U. alpina | Schweigert et al., 2003, Plate 10, Figure 2 | - | 3.69 | - | 4.69 | 1.80 | 2.93 | - | 1.71 | 0.82 | 1.61 | 2.38 | 1.39 | 0.51 | 1.96 | - | - | 0.76 |
MNSM no. i10863 | U. alpina | Schweigert et al., 2003, Plate 10, Figure 3 | - | 5.29 | - | 5.40 | 2.46 | 4.06 | - | 2.83 | 1.44 | 1.97 | 2.92 | 1.03 | 0.73 | 1.37 | - | - | 0.84 |
MNSM no. i10851 part | U. alpina | Schweigert et al., 2003, Plate 10, Figure 5 | - | 4.20 | - | 6.00 | 2.70 | 4.16 | - | 2.82 | 1.24 | 1.56 | 2.89 | 1.02 | 0.79 | 1.26 | - | - | 0.93 |
MNSM no. i10851 counterpart | U. alpina | Schweigert et al., 2003, Plate 10, Figure 5 | - | 4.30 | - | 6.50 | 2.31 | 4.28 | - | 2.97 | 1.21 | 1.33 | 3.02 | 1.02 | 0.91 | 1.10 | - | - | 0.76 |
APPENDIX 3
R Code for Boxplots and Wilcoxon-Ranked Sum tests (R Studio)
##### BOX PLOTS###
# boxes colored by species
#index ratio
boxplot(Appendix2Rtrim$`IndexLengthtoHeight` ~ as.factor(Appendix2Rtrim$Species), main="Index length to index height ratio in Appendix2RTRIMMED Uncinids", xlab="Species", col=c('steelblue3', 'darkorchid3', 'dark green', 'dark red'))
#notch ratio
boxplot(Appendix2Rtrim$`NotchLengthtoNotchHeight` ~ as.factor(Appendix2Rtrim$Species), main="Notch length to notch height ratio in Appendix2RTRIMMED Uncinidas", xlab="Species", col=c('steelblue3', 'darkorchid3', 'dark green', 'dark red'))
#Length
boxplot(Appendix2Rtrim$`PropodusLength` ~ as.factor(Appendix2Rtrim$Species), main="Length of chelae in Appendix2RTRIMMED Uncinids", xlab="Species", col=c('steelblue3', 'darkorchid3', 'dark green', 'dark red'))
#Manus Length:Index Length
boxplot(Appendix2Rtrim$`ManustoIndexLength` ~ as.factor(Appendix2Rtrim$Species), main="Manus length to index length of Appendix2RTRIMMED Uncinids", xlab="Species", col=c('steelblue3', 'darkorchid3', 'dark green', ' dark red'))
#Dactylus Thickness:Index Height
boxplot(Appendix2Rtrim$`DactylusThicknesstoIndexHeight` ~ as.factor(Appendix2Rtrim$Species), main="Dactylus Thickness to index height of Appendix2RTRIMMED Uncinids", xlab="Species",
col=c('steelblue3', 'darkorchid3', 'dark green', ' dark red'))
#Manus Height Min:Max
boxplot(Appendix2Rtrim$`ManusHeightAttachtoManusHeight` ~ as.factor(Appendix2Rtrim$Species), main="Manus height at attachment to Max height to index length of Appendix2RTRIMMED Uncinids", xlab="Species", col=c('steelblue3', 'darkorchid3', 'dark green', ' dark red'))
###### Species Comparisons ######
UPAC <- subset(Appendix2Rtrim, Species=="U. pacifica",)
UPOS <- subset(Appendix2Rtrim, Species=="U. posidoniae",)
UALP <- subset(Appendix2Rtrim, Species=="U. alpina",)
UOLL <- subset(Appendix2Rtrim, Species=="U. ollerenshawi",)
##### TABLE 3 #####
#p-value results of the Wilcoxon Ranked-Sum tests comparing dactylus thickness to index height of each uncinid species. This test was employed to distinguish the differences in the depth of the index of each species. Alpha value = 0.05; see Table 2 for measurements and Appendix 3 for R code
#alternative hypothesis: true location shift is not equal to 0
#DactylusThicknesstoIndexHeight#
PASVPOS <- wilcox.test(na.omit(UPAC$DactylusThicknesstoIndexHeight), na.omit(UPOS$DactylusThicknesstoIndexHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVPOS
#data: na.omit(UPAC$DactylusThicknesstoIndexHeight) and na.omit(UPOS$DactylusThicknesstoIndexHeight)
#W = 3, p-value = 0.0001852
PASVALP <- wilcox.test(na.omit(UPAC$DactylusThicknesstoIndexHeight), na.omit(UALP$DactylusThicknesstoIndexHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVALP
#data: na.omit(UPAC$DactylusThicknesstoIndexHeight) and na.omit(UALP$DactylusThicknesstoIndexHeight)
#W = 2, p-value = 0.006216
PASVOLL <- wilcox.test(na.omit(UPAC$DactylusThicknesstoIndexHeight), na.omit(UOLL$DactylusThicknesstoIndexHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVOLL
#na.omit(UPAC$DactylusThicknesstoIndexHeight) and na.omit(UOLL$DactylusThicknesstoIndexHeight)
#W = 2, p-value = 0.006216
OLLVPOS <- wilcox.test(na.omit(UOLL$DactylusThicknesstoIndexHeight), na.omit(UPOS$DactylusThicknesstoIndexHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVPOS
#data: na.omit(UOLL$DactylusThicknesstoIndexHeight) and na.omit(UPOS$DactylusThicknesstoIndexHeight)
#W = 17, p-value = 0.2674
OLLVALP <- wilcox.test(na.omit(UOLL$DactylusThicknesstoIndexHeight), na.omit(UALP$DactylusThicknesstoIndexHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVALP
#data: na.omit(UOLL$DactylusThicknesstoIndexHeight) and na.omit(UALP$DactylusThicknesstoIndexHeight)
#W = 10, p-value = 0.6905
ALPVPOS <- wilcox.test(na.omit(UALP$DactylusThicknesstoIndexHeight), na.omit(UPOS$DactylusThicknesstoIndexHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
ALPVPOS
#data: na.omit(UALP$DactylusThicknesstoIndexHeight) and na.omit(UPOS$DactylusThicknesstoIndexHeight)
#W = 24, p-value = 0.7427
##### TABLE 4 #####
#p-value results of the Wilcoxon Ranked-Sum tests comparing the index length to index height of each uncinid species. This test was employed to determine the differences in the shape of the index between species. Alpha value = 0.05; see Table 2 for measurements and Appendix 3 for R code.
#alternative hypothesis: true location shift is not equal to 0
#IndexLengthtoHeight#
PASVPOS <- wilcox.test(na.omit(UPAC$IndexLengthtoHeight), na.omit(UPOS$IndexLengthtoHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVPOS
#data: na.omit(UPAC$IndexLengthtoHeight) and na.omit(UPOS$IndexLengthtoHeight)
#W = 143, p-value = 8.012e-07
PASVALP <- wilcox.test(na.omit(UPAC$IndexLengthtoHeight), na.omit(UALP$IndexLengthtoHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVALP
#data: na.omit(UPAC$DactylusThicknesstoIndexHeight) and na.omit(UALP$DactylusThicknesstoIndexHeight)
#W = 30, p-value = 0.849
PASVOLL <- wilcox.test(na.omit(UPAC$IndexLengthtoHeight), na.omit(UOLL$IndexLengthtoHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVOLL
#na.omit(UPAC$IndexLengthtoHeight) and na.omit(UOLL$IndexLengthtoHeight)
#W = 65, p-value = 0.0002334
OLLVPOS <- wilcox.test(na.omit(UOLL$IndexLengthtoHeight), na.omit(UPOS$IndexLengthtoHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVPOS
#data: na.omit(UOLL$IndexLengthtoHeight) and na.omit(UPOS$IndexLengthtoHeight)
#W = 14, p-value = 0.1451
OLLVALP <- wilcox.test(na.omit(UOLL$IndexLengthtoHeight), na.omit(UALP$IndexLengthtoHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVALP
#data: na.omit(UOLL$IndexLengthtoHeight) and na.omit(UALP$IndexLengthtoHeight)
#W = 0, p-value = 0.007937
ALPVPOS <- wilcox.test(na.omit(UALP$IndexLengthtoHeight), na.omit(UPOS$IndexLengthtoHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
ALPVPOS
#data: na.omit(UALP$IndexLengthtoHeight) and na.omit(UPOS$IndexLengthtoHeight)
#W = 55, p-value = 0.0004579
##### TABLE 5 #####
#. p-value results of the Wilcoxon Ranked-Sum tests comparing the manus height at its minimum and its maximum for each uncinid species. This test was employed to determine the differences in the overall shape of the claw between species. Alpha value = 0.05; see Table 2 for measurements and Appendix 3 for R code.
#alternative hypothesis: true location shift is not equal to 0
#ManusHeightAttachtoManusHeight#
PASVPOS <- wilcox.test(na.omit(UPAC$ManusHeightAttachtoManusHeight), na.omit(UPOS$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVPOS
#data: na.omit(UPAC$ManusHeightAttachtoManusHeight) and na.omit(UPOS$ManusHeightAttachtoManusHeight)
#W = 1, p-value = 1.134e-05
PASVALP <- wilcox.test(na.omit(UPAC$ManusHeightAttachtoManusHeight), na.omit(UALP$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVALP
#data: na.omit(UPAC$ManusHeightAttachtoManusHeight) and na.omit(UALP$ManusHeightAttachtoManusHeight)
#W = 0, p-value = 0.1818
PASVOLL <- wilcox.test(na.omit(UPAC$ManusHeightAttachtoManusHeight), na.omit(UOLL$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVOLL
#na.omit(UPAC$ManusHeightAttachtoManusHeight) and na.omit(UOLL$ManusHeightAttachtoManusHeight)
#W = 1, p-value = 0.001332
OLLVPOS <- wilcox.test(na.omit(UOLL$ManusHeightAttachtoManusHeight), na.omit(UPOS$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVPOS
#data: na.omit(UOLL$ManusHeightAttachtoManusHeight) and na.omit(UPOS$ManusHeightAttachtoManusHeight)
#W = 19, p-value = 0.3773
OLLVALP <- wilcox.test(na.omit(UOLL$ManusHeightAttachtoManusHeight), na.omit(UALP$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVALP
#data: na.omit(UOLL$ManusHeightAttachtoManusHeight) and na.omit(UALP$ManusHeightAttachtoManusHeight)
#W = 0, p-value = 0.3333
ALPVPOS <- wilcox.test(na.omit(UALP$ManusHeightAttachtoManusHeight), na.omit(UPOS$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
ALPVPOS
#data: na.omit(UALP$ManusHeightAttachtoManusHeight) and na.omit(UPOS$ManusHeightAttachtoManusHeight)
#W = 11, p-value = 0.1667
##### TABLE 6 #####
#p-value results of the Wilcoxon Ranked-Sum tests comparing the manus length to index length of each uncinid species. This test was employed to determine the differences in the relative length of the fingers between species. Alpha value = 0.05; see Table 2 for measurements and Appendix 3 for R code.
#alternative hypothesis: true location shift is not equal to 0
#ManustoIndexLength#
PASVPOS <- wilcox.test(na.omit(UPAC$ManustoIndexLength), na.omit(UPOS$ManustoIndexLength), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVPOS
#data: na.omit(UPAC$ManustoIndexLength) and na.omit(UPOS$ManustoIndexLength)
#W = 43, p-value = 0.4262
PASVALP <- wilcox.test(na.omit(UPAC$ManustoIndexLength), na.omit(UALP$ManustoIndexLength), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVALP
#data: na.omit(UPAC$ManustoIndexLength) and na.omit(UALP$ManustoIndexLength)
#W = 0, p-value = 0.1818
PASVOLL <- wilcox.test(na.omit(UPAC$ManustoIndexLength), na.omit(UOLL$ManustoIndexLength), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVOLL
#na.omit(UPAC$ManustoIndexLength) and na.omit(UOLL$ManustoIndexLength)
#W = 34, p-value = 0.3097
OLLVPOS <- wilcox.test(na.omit(UOLL$ManustoIndexLength), na.omit(UPOS$ManustoIndexLength), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVPOS
#data: na.omit(UOLL$ManustoIndexLength) and na.omit(UPOS$ManustoIndexLength)
#W = 21, p-value = 0.5096
OLLVALP <- wilcox.test(na.omit(UOLL$ManustoIndexLength), na.omit(UALP$ManustoIndexLength), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVALP
#data: na.omit(UOLL$ManusHeightAttachtoManusHeight) and na.omit(UALP$ManusHeightAttachtoManusHeight)
#W = 0, p-value = 0.3333
ALPVPOS <- wilcox.test(na.omit(UALP$ManustoIndexLength), na.omit(UPOS$ManustoIndexLength), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
ALPVPOS
#data: na.omit(UALP$ManustoIndexLength) and na.omit(UPOS$ManustoIndexLength)
#W = 11, p-value = 0.1667
##### TABLE 5 #####
#. p-value results of the Wilcoxon Ranked-Sum tests comparing the manus height at its minimum and its maximum for each uncinid species. This test was employed to determine the differences in the overall shape of the claw between species. Alpha value = 0.05; see Table 2 for measurements and Appendix 3 for R code.
#alternative hypothesis: true location shift is not equal to 0
#ManusHeightAttachtoManusHeight#
PASVPOS <- wilcox.test(na.omit(UPAC$ManusHeightAttachtoManusHeight), na.omit(UPOS$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVPOS
#data: na.omit(UPAC$ManusHeightAttachtoManusHeight) and na.omit(UPOS$ManusHeightAttachtoManusHeight)
#W = 1, p-value = 1.134e-05
PASVALP <- wilcox.test(na.omit(UPAC$ManusHeightAttachtoManusHeight), na.omit(UALP$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVALP
#data: na.omit(UPAC$ManusHeightAttachtoManusHeight) and na.omit(UALP$ManusHeightAttachtoManusHeight)
#W = 0, p-value = 0.1818
PASVOLL <- wilcox.test(na.omit(UPAC$ManusHeightAttachtoManusHeight), na.omit(UOLL$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVOLL
#na.omit(UPAC$ManusHeightAttachtoManusHeight) and na.omit(UOLL$ManusHeightAttachtoManusHeight)
#W = 1, p-value = 0.001332
OLLVPOS <- wilcox.test(na.omit(UOLL$ManusHeightAttachtoManusHeight), na.omit(UPOS$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVPOS
#data: na.omit(UOLL$ManusHeightAttachtoManusHeight) and na.omit(UPOS$ManusHeightAttachtoManusHeight)
#W = 19, p-value = 0.3773
OLLVALP <- wilcox.test(na.omit(UOLL$ManusHeightAttachtoManusHeight), na.omit(UALP$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVALP
#data: na.omit(UOLL$ManusHeightAttachtoManusHeight) and na.omit(UALP$ManusHeightAttachtoManusHeight)
#W = 0, p-value = 0.3333
ALPVPOS <- wilcox.test(na.omit(UALP$ManusHeightAttachtoManusHeight), na.omit(UPOS$ManusHeightAttachtoManusHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
ALPVPOS
#data: na.omit(UALP$ManusHeightAttachtoManusHeight) and na.omit(UPOS$ManusHeightAttachtoManusHeight)
#W = 11, p-value = 0.1667
##### TABLE 7 #####
#p-value results of the Wilcoxon Ranked-Sum tests comparing the notch length to the notch height of each uncinid species. Alpha value = 0.05; see Table 2 for measurements and Appendix 3 for R code.
#alternative hypothesis: true location shift is not equal to 0
#NotchLengthtoNotchHeight#
PASVPOS <- wilcox.test(na.omit(UPAC$NotchLengthtoNotchHeight), na.omit(UPOS$NotchLengthtoNotchHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVPOS
#data: na.omit(UPAC$NotchLengthtoNotchHeight) and na.omit(UPOS$NotchLengthtoNotchHeight)
#W = 38, p-value = 0.588
PASVALP <- wilcox.test(na.omit(UPAC$NotchLengthtoNotchHeight), na.omit(UALP$NotchLengthtoNotchHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVALP
#data: na.omit(UPAC$NotchLengthtoNotchHeight) and na.omit(UALP$NotchLengthtoNotchHeight)
#W = 44, p-value = 0.2887
PASVOLL <- wilcox.test(na.omit(UPAC$NotchLengthtoNotchHeight), na.omit(UOLL$NotchLengthtoNotchHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
PASVOLL
#data: na.omit(UPAC$NotchLengthtoNotchHeight) and na.omit(UOLL$NotchLengthtoNotchHeight)
#W = 21, p-value = 0.1274
OLLVPOS <- wilcox.test(na.omit(UOLL$NotchLengthtoNotchHeight), na.omit(UPOS$NotchLengthtoNotchHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVPOS
#data: na.omit(UOLL$NotchLengthtoNotchHeight) and na.omit(UPOS$NotchLengthtoNotchHeight)
#W = 31, p-value = 0.1807
OLLVALP <- wilcox.test(na.omit(UOLL$NotchLengthtoNotchHeight), na.omit(UALP$NotchLengthtoNotchHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
OLLVALP
#na.omit(UOLL$NotchLengthtoNotchHeight) and na.omit(UALP$NotchLengthtoNotchHeight)
#W = 27, p-value = 0.0303
ALPVPOS <- wilcox.test(na.omit(UALP$NotchLengthtoNotchHeight), na.omit(UPOS$NotchLengthtoNotchHeight), alternative = c("two.sided", "less", "greater"), conf.level = 0.99, paired=F)
ALPVPOS
#data: na.omit(UALP$NotchLengthtoNotchHeight) and na.omit(UPOS$NotchLengthtoNotchHeight)
#W = 7, p-value = 0.1061
APPENDIX 4
Photographs of all Uncinidae specimens used in this study (whitened with ammonium chloride unless otherwise noted). Descriptions of photograph techniques can be found in the methods.
FIGURE A4-1. Dactylus of Uncina pacifica, specimen TMP 2002.043.0002 (scale is in mm, numbers are cm).
FIGURE A4-2. Three chelae, two Uncina pacifica and one Uncina ollerenshawi chelae, specimen TMP 2002.043.0003 (scale is in mm, numbers are cm).
FIGURE A4-3. Articulated chela of Uncina pacifica, specimen TMP 2002.043.0004 (scale is in mm, numbers are cm).
FIGURE A4-4. Part and counterpart, fixed finger, and partial hand of Uncina pacifica, specimen TMP 2002.043.0006 (scale is in mm, numbers are cm).
FIGURE A4-5. Dactylus of Uncina pacifica, specimen TMP 2002.043.0007 (scale is in mm, numbers are cm).
FIGURE A4-6. Fixed finger and hand of Uncina pacifica, specimen TMP 2002.043.0008 (scale is in mm, numbers are cm).
FIGURE A4-7. Dactylus of Uncina pacifica, specimen TMP 2002.043.0012 (scale is in mm, numbers are cm).
FIGURE A4-8. Fixed finger and hand of Uncina pacifica, specimen TMP 2002.043.0015 (scale is in mm, numbers are cm).
FIGURE A4-9. Major chela (upper left) and dactylus (lower right) of Uncina pacifica, specimen TMP 2002.043.0019 (scale is in mm, numbers are cm).
FIGURE A4-10. Major chela of Uncina pacifica (part and counterpart), specimen TMP 2014.021.0003 (scale is in mm, numbers are cm).
FIGURE A4-11. Minor chela of Uncina pacifica, specimen TMP 2014.021.0004 (scale is in mm, numbers are cm).
FIGURE A4-12. Minor chela of Uncina pacifica, specimen TMP 2014.021.0007 (scale is in mm, numbers are cm).
FIGURE A4-13. Dactylus of Uncina pacifica, specimen TMP 2014.021.0033 (scale is in mm, numbers are cm).
FIGURE A4-14. Hand and fixed finger of Uncina pacifica (part and counterpart), specimen TMP 2014.021.0034 (scale is in mm, numbers are cm).
FIGURE A4-15. Body molt of Uncina ollerenshawi, specimen TMP 2014.021.0052 (scale is in mm, numbers are cm).
FIGURE A4-16. Dactylus of Uncina pacifica, specimen TMP 2015.050.0038 (scale is in mm, numbers are cm).
FIGURE A4-17. Minor claw of Uncina pacifica, specimen TMP 2015.050.0039 (scale is in mm, numbers are cm).
FIGURE A4-18. Dactylus of Uncina pacifica, specimen TMP 2015.050.0069 (scale is in mm, numbers are cm).
FIGURE A4-19. Partial fixed finger of Uncina pacifica, specimen TMP 2015.050.0069 (scale is in mm, numbers are cm).
FIGURE A4-20. Uncina ollerenshawi chela with the carpus and partial merus articulated, specimen TMP 2015.051.0063 (scale is in mm, numbers are cm).
FIGURE A4-21. Partial major chela of Uncina pacifica, merus, carpus and partial ischium included, specimen TMP 2016.027.0003 (scale is in mm, numbers are cm).
FIGURE A4-22. Body fossil of Uncina pacifica, major and minor chelae present, specimen TMP 2017.015.0001 (scale is in mm, numbers are cm).
FIGURE A4-23. Minor chela of Uncina pacifica, specimen TMP 2017.015.0003 (scale is in mm, numbers are cm).
FIGURE A4-24. Chela of Uncina pacifica, specimen TMP 2017.015.0005 (scale is in mm, numbers are cm).
FIGURE A4-25. Minor chela (fixed finger) of Uncina pacifica, specimen TMP 2017.015.0007 (scale is in mm, numbers are cm).
FIGURE A4-26. Dactylus of Uncina pacifica, specimen TMP 2018.024.0005 (scale is in mm, sample not whitened).
FIGURE A4-27. Major chela of Uncina pacifica, specimen TMP 2018.024.0003 (scale is in mm, numbers are cm).
FIGURE A4-28. Partial minor chela (fixed finger) of Uncina ollerenshawi, specimen TMP 2018.024.0006 (scale is in mm, numbers are cm).
FIGURE A4-29. Partial minor chela (fixed finger) of Uncina ollerenshawi, specimen TMP 2018.024.0007 (scale is in mm, numbers are cm).
FIGURE A4-30. Dactylus of Uncina ollerenshawi, specimen TMP 2018.024.0009 (scale is in mm, numbers are cm).
FIGURE A4-31. Partial minor chela (fixed finger) of Uncina pacifica (partially obscured), specimen TMP 2018.024.0035 (scale is in mm, numbers are cm).
FIGURE A4-32. Body fossil of the Uncina ollerenshawi Holotype, specimen GSC 80067 (scale is in mm, numbers are cm).
FIGURE A4-33. Dactylus of Uncina ollerenshawi, specimen TMP 2016.027.0008 (scale is in mm). Specimen not whitened.
FIGURE A4-34. Partial minor chela (fixed finger) of Uncina pacifica, specimen TMP 2018.024.0004 (scale is in mm, numbers are cm). Specimen not whitened.
FIGURE A4-35. Major chela of Uncina pacifica, specimen TMP 2002.043.0005 (scale is in mm, numbers are cm). Specimen not whitened.
FIGURE A4-36. Major chela of Uncina ollerenshawi, specimen TMP 2013.036.0002 (scale is in mm, numbers are cm). Specimen not whitened.
FIGURE A4-37. Major chela of Uncina pacifica, specimen split across two samples, TMP 2013.0036.0004 and TMP 2014.021.0002 (scale is in mm, numbers are cm). Specimen not whitened.
FIGURE A4-38. Partial dactylus of Uncina pacifica, specimen TMP 2014.021.0008 (scale is in cm). Specimen not whitened.
FIGURE A4-39. Major chela of Uncina pacifica, specimen TMP 2014.021.0023 (part and counterpart) (scale is in cm). Specimen not whitened.
FIGURE A4-40. Major chela of Uncina pacifica, specimen TMP 2015.050.0011 (part and counterpart) (scale is in mm). Specimen not whitened.
FIGURE A4-41. Major chelae of Uncina posidoniae, specimen MB.A.3826 (scale is in cm). Specimen not whitened.
Note: Upon completion of the review process of this manuscript, it was brought to our attention that another specimen of Uncina ollerenshawi existed (TMP 2005.028.0004). The specimen, collected by Russell Hall from the East Tributary of Ya Ha Tinda, was sent to the Staatliches Museum für Naturkunde, Stuttgart, Germany, and subsequently not available for study in this work.
FIGURE A4-42. Body fossil of Uncina ollerenshawi (part and counterpart), specimen TMP 2005.028.0004 (scale closest to specimen is in mm, numbers are cm). Specimen not whitened.