TABLE 1. Measurements, body proportions, and meristic data of the Amakusaichthys benammii sp. nov. specimens. All measurements are in millimeters. The proportions correspond to the % of the LS; “-” shows unavailable data.
Measurement | IGM 14027 |
IGM 14028 |
IGM 14029 |
IGM 14030 |
IGM 14031 |
IGM 14032 |
IGM 14034 |
Means or modes |
Total length (TL) | 454 | 473 | - | - | - | - | - | - |
Standard length (SL) | 391 | 385 | 310+ | - | 215+ | - | - | 388 |
Head length (HL) HL as % of SL |
70 17.9 |
72 18.7 |
67 - |
- - |
72 - |
89 - |
- | 74.2 18.2 |
Head height (HH) HH as % of SL |
52 13.3 |
51 13.2 |
50 - |
- - |
57? - |
62? - |
- | 51 13.2 |
Maximum body height (MBH) MBH as % of SL |
54 13.8 |
55 14.2 |
52? - |
- - |
- - |
- - |
- | 54.5 14 |
Prepelvic length (PPL) PPL as % of SL |
274 70 |
270 70 |
227? - |
- - |
180? | - - |
- | 272 70 |
Predorsal length (PDL) HL as % of SL |
? (-) |
310 80.5 |
- - |
- - |
- - |
- - |
- | 310 80.5 |
Dorsal fin length (DFL) DFL as % of SL |
- - |
21 5.4 |
- - |
- - |
- - |
- - |
- | 21 5.4 |
Preanal length (PAL) PAL as % of SL |
323 (82.6) |
319 81 |
- - |
- - |
- - |
- - |
- | 314 81 |
Anal fin length (AFL) AFL as % of SL |
24? (6.1) |
35 9 |
- | - - |
- - |
- - |
- | 31 9 |
Caudal peduncle height (CPH) CPH as % of SL |
21 (5.3) |
20 5.1 |
- - |
- - |
- - |
- - |
- - |
20.2 5.2 |
Total vertebrae | 78 | 78 | 65+ | 58+ | 42+ | - | 73+ | 78 |
Abdominal vertebrae | 54 | 54 | - | - | - | - | 54? | 54 |
Caudal vertebrae | 22 | 22 | - | - | - | - | 19? | 22 |
Pectoral fin rays | 12 | 12 | 12 | 11? | 12 | 12 | - | 12 |
Pelvic fin rays | 8 | 8 | 7+ | - | 6+ | - | - | 8 |
Dorsal fin rays | 4+ | 13 | - | - | - | - | - | 13 |
Anal rays | 6+ | 11 | - | - | - | - | - | 11 |
TABLE 2. Nominal valid species described or referred to the order Ichthyodectiformes.
Species | Age. Distribution. |
Africathrissops weileri Taverne, 2010 | Aptian. Gabon |
Aidachar pankowskii (Forey and Cavin, 2007) | Cenomanian. Morocco |
Aidachar paludalis Nessov, 1981 (see Mkhitaryan and Averianov, 2011) | Turonian. Uzbekistan |
Allothrissops mesogaster (Agassiz, 1833) | Early Tithonian. Germany |
Allothrissops regleyi (Thiollière, 1854) | Kimmeridgian. Francia |
Allothrissops salmoneus (Blainville, 1818) | Kimmeridgian. Europe |
Altamuraichthys meleleoi Taverne, 2016 | Campanian-Maastrichtian. Italy |
Amakusaichthys benammii Alvarado-Ortega, this work | Campanian, Mexico |
Amakusaichthys goshouraensis Yabumoto, Hirose, and Brito, 2018 | Santonian. Japan |
Antartithrissops australis Arratia, Scasso, and Kiessling, 2004 | Tithonian. Antarctica |
Ascalabothrissops voelkli Arratia, 2000 | Kimmeridgian. Germany |
Bardackichthys carteri Hacker and Shimada, 2021 | Cenomanian. USA |
Capassoichthys alfonsoi Taverne, 2015 | Campanian-Maastrichtian. Italy |
Chirocentrites coroninii Heckel, 1849 | Cenomanian. Slovenia |
Chirocentrites neocomiensis (Pictet, 1858) | Hauterivian. France |
Chirocentrites ? calvolini Costa, 1865 | Albian. Italy |
Chiromystus alagoensis Jordan, 1910 | Early Cretaceous, Brazil |
Chiromystus guinensis (Weiler, 1922) | Aptian. Africa |
Chiromystus mawsoni Cope, 1885 | HauterivianAlbian. Brazil |
Chuhsiungichthys japonicus Yabumoto, 1994 | Hauterivian-Barremian. Japan |
Chuhsiungichthys tsanglingensis Lew, 1974 | Early Cretaceous. China |
Chuhsiungichthys yanagidai Yabumoto, 1994 | Hauterivian-Barremian. Japan |
Cladocyclus gardneri, Agassiz, 1841 | Aptian. South America |
Cladocyclus geddesi Berrell, Alvarado-Ortega, Yabumoto, and Salisbury, 2014 | Albian. Australia |
Cladocynodon araripensis Mayrinck, Ribeiro, Assine, and Spigolon, 2023 | Aptian. Brazil |
Cooyoo australis (Woodward, 1894) | Albian. Australia |
Dugaldia emmilta Cavin and Berrell, 2019 | Albian. Australia |
Eubiodectes libanicus (Pictet and Humbert, 1866) | Cenomanian. Lebanon |
Faugichthys loryi Taverne and Chanet, 2000 | Albian. France |
Furloichthys bonarellii Taverne and Capasso, 2018 | Cenomanian. Italy |
Garganoichthys alfonsoi Taverne, 2009 | Santonian. Italy |
Gillicus arcuatus Cope, 1875 | Albian-Campanian. North America |
Gillicus serridens Woodward, 1901 | Albian. England |
Ghrisichthys bardacki (Cavin, 1997) | Turonian. Morocco |
Gwawinapterus beardi Arbour, Currie, 2011 | Late Cretaceous. Canada |
Heckelichthys preopercularis Baños-Rodríguez, González-Rodríguez, Wilson, and González-Martínez, 2020 | Albian-Cenomanian limit. Mexico |
Heckelichthys microdon (Heckel, 1849) | Barremian-Cenomanian. Europe |
Heckelichthys vexillifer (Heckel, 1856) | Neocomian. Morroco |
Ichthyodectes ctenodon (Leidy, 1857) | Albian-Campanian. USA |
Ichthyodectes minor Dixon, 1850 | Albian-Campanian. Europe |
Itaparica woodwardi (Silva-Santos, 1949) | Hauterivian-Albian. Brazil |
Jinjuichthys cheongi Kim, Chang, Wu, and Kim, 2014 | Albian. Japan |
Mesoclupea showchangensis Ping and Yen, 1933 | Early Cretaceous. China |
Occithrissops willsoni Schaeffer and Patterson, 1984 | Bathonian. USA |
Ogunichthys triangularis Alvarado-Ortega and Brito 2010 | Aptian?. Brazil |
Pachythrissops laevis Woodward, 1919 | Purbeckian. England |
Pachythrissops propterus (Wagner, 1863) | Tithonian. Germany |
Postredectes harranaensis Kaddumi, 2009 | Maastrichtian-Paleocene?, Jordan |
Proportheus kameruni Jaekel, 1909 | Turonian-Cenomanian. Africa |
Prosaurodon pygmeus (Loomis, 1900) (see Stewart, 1999) | Santonian-Campanian. USA |
Prymnetes longiventer Cope, 1871 | Early Cretaceous?. Mexico |
Saurocephalus lanciformis Harlan, 1824 | Turonian-Maastrichtian. America |
Saurocephalus woodwardi Davies, 1878 | Upper Cretaceous. Europe |
Saurodon elongatus Taverne and Bronzi 1999 | Campanian-Maastrichtian. Italy |
Saurodon intermedius Newton, 1878 | Cenomanian. England |
Saurodon leanus Hays, 1830 | Cenomanian-Campanian. North America |
Sultanuvaisia antiqua Nessov 1981 | Late Cretaceous. Kazakhstan |
Thrissops cephalus Agassiz, 1844 | Oxfordian. England |
Thrissops cirinensis Nybelin, 1964 (see Taverne, 1977) | Kimmeridgian. Germany |
Thrissops costalis Egerton, 1845 | Oxfordian. England |
Thrissops curtus Woodward, 1919 | Portlandian. England |
Thrissops exiguus Bassani, 1879 | Cenomanian. Slovenia |
Thrissops formosus Agassiz, 1833 (see Taverne, 1977) | Kimmeridgian. Germany |
Thrissops gracilis Heckel, 1849 | Cenomanian. Slovenia |
Thrissops portlandicus Woodward, 1895 | Portlandian. England |
Thrissops rochei Sauvage, 1893 | Kimmeridgian. France |
Thrissops subovatus Münster (in Agassiz, 1844) (see Taverne, 1977) | Kimmeridgian. Germany |
Thrissops volganensis Koslov, 1928 | Portlandian. Russia |
Unamichthys espinosai Alvarado-Ortega, 2004 | Albian. Mexico |
Vallecillichthys multivertebratum Blanco and Cavin, 2003 | Turonian. Mexico |
Verraesichthys bloti Taverne, 2010 | Aptian. Africa |
Xiphactinus audax Leidy, 1870 | Cenomanian-Campanian. North America |
Xiphactinus gaultinus Newton, 1877 | Albian-Cenomanian. England |
Xiphactinus mantelli Newton, 1877 | Cenomanian-Campanian. Europe |
Xiphactinus vetus (Leidy, 1870) | Campanian-Maastrichtian. USA |