FIGURE 1. Gracanicanetta happi gen. et sp. nov., holotype, specimen ISEA AF/GRA1 from Gračanica, Bosnia and Herzegovina, Early-Middle Miocene. A, slab after exposing all the bones and removing the os carpi ulnare; B, fragment of slab before removal of the os carpi ulnare. Abbreviations: gas, gastroliths; lca, left carpometacarpus; lcl, left clavicula (omal fragment); lco, left coracoid; lcr, left os carpi radiale; lcu, left os carpi ulnare; lhu, left humerus; lpp, left phalanx proximalis digiti majoris; lra, left radius; lul, left ulna; man, mandibula; pel, pelvis; rcl, right clavicula (omal fragment); rco, right coracoid; rfe, right femur; rhu, right humerus; rra, right radius; rsc, right scapula; rul, right ulna; sku, skull; ste, sternum.
FIGURE 2. Comparison of the skull and mandible. A, Gracanicanetta happi gen. et sp. nov., holotype, specimen ISEA AF/GRA1 from Gračanica, Bosnia and Herzegovina, Early-Middle Miocene, in ventrolateral view; B, Clangula hyemalis (Mergini) in dorsal view; C, Spatula querquedula (Anatini) in dorsal view. Abbreviations: fne, foramina neurovascularia; lrm, left ramus mandibulae; rrm, right ramus mandibulae; pme, procesus medialis; por, preorbital region; ppo, procesus postorbitalis; pre, procesus retroarticularis.
FIGURE 3. A, sternum and pelvis of Gracanicanetta happi gen. et sp. nov., holotype, specimen ISEA AF/GRA1 from Gračanica, Bosnia and Herzegovina, Early-Middle Miocene, in dorsal view; B, sternum of Bucephala clangula (Mergini) in dorsal view; C, sternum of Spatula querquedula (Anatini) in dorsal view. The extant species are shown in the same size as the fossil to facilitate comparison. Abbreviations: gas, gastroliths; ila, incisura lateralis; mco, margo costalis; pco, pila coracoidea; pel, pelvis; rfe, right femur; ste, sternum.
FIGURE 4. Gracanicanetta happi gen. et sp. nov., holotype, specimen ISEA AF/GRA1 from Gračanica, Bosnia and Herzegovina, Early-Middle Miocene. A, right coracoid in dorsal view, right scapula in ventrolateral view, and omal fragment of right clavicula in lateral view; B, computed tomography scan of right coracoid in ventral view; C, omal fragment of right coracoid in dorsomedial view. Abbreviations: csc, cotyla scapularis; fac, facies articularis clavicularis; fah, facies articularis humeralis; pac, procesus acrocoracoideus; rcl, right clavicula (omal fragment); rco, right coracoid; rsc, right scapula; smc, sulcus m. coracoidei; ssa, shelf-shaped attachment.
FIGURE 5. Gracanicanetta happi gen. et sp. nov., holotype, specimen ISEA AF/GRA1 from Gračanica, Bosnia and Herzegovina, Early-Middle Miocene. A, wing bones; B, computed tomography scan of left carpometacarpus in dorsal view; C, proximal humerus of Protomelanitta gracilis, redrawn from Zelenkov (2017: fig.1c); D, schematic drawing of the proximal humerus of Gracanicanetta happi. Abbreviations: cbi, crista bicipitalis (possible shape); cde, crista deltopectoralis; chu, caput humeri; csr, capital shaft ridge; fbr, fossa brachialis; fin, fossa infratrochlearis; fpd, fossa pneumotricipitalis dorsalis; fpv, fossa pneumotricipitalis ventralis; lca, left carpometacarpus; lcr, left os carpi radiale; lhu, left humerus; lpp, left phalanx proximalis digiti majoris; lra, left radius; lul, left ulna; nic, notch in incisura capitis; ntc, notch in caudal rim of trochlea carpalis; pex, processus extensorius; pfl, processus flexorius; ppi, processus pisiformis; rhu, right humerus; rra, right radius; rul, right ulna; smd, synostosis metacarpalis distalis; tdo, tuberculum dorsale; tve, tuberculum ventrale; ved, ventral edge of the condylus dorsalis.
FIGURE 6. Comparison of the os carpi ulnare. A-B, Gracanicanetta happi gen. et sp. nov., holotype, specimen ISEA AF/GRA1 from Gračanica, Bosnia and Herzegovina, Early-Middle Miocene; C-D, Aythya ferina (Aythyini); E-F, Spatula querquedula (Anatini); G-H, Bucephala clangula (Mergini). Specimens in the left column (A, C, E and G) are shown in distal view, and those in the right column (B, D, F and H) in dorsal view. The extant species are shown to be the same size as the fossil to facilitate comparison. Abbreviations: cbr, crista breve; clo, crista longum.
