TABLE 1. Known species of Brachymetopus (Acutimetopus) Hahn and Hahn, 1985. Permian species are highlighted in bold. Here and below is a list of species according to their year of description.
| Species | Part | Stratigraphy | Geography |
| B. (A.) gracilis Heritsch, 1931 | Cph | Upper Pennsylvanian, Stephanian (~Kasimovian-Gzhelian) | Austria, Carinthia |
| B. (A.) moelleri Weber, 1932 | Cph | Cisuralian, Asselian | Russia, Perm Krai |
| B. (A.) sp. Weber, 1937 | Cph | Middle Pennsylvanian, Moscovian | Donetsk Basin, Krasnaya Mogila railway station |
| B. (A.) caucasicus Licharew in Weber, 1944 | Py | Lopingian, Changhsingian | Russia, Krasnodar Krai, Malaya Laba River Basin |
| B. (A.) kansasensis (Weller, 1944) type species | Cc | Upper Pennsylvanian (U. Pennsylvanian), upper part of the Haskell Limestone | USA, Kansas, Leavenworth County |
| B. (A.) pseudometopina Gauri et Ramovš, 1964 | Cph, Py | Upper Pennsylvanian, Gzhelian | Slovenia, Southern Karavanke |
| B. (A.) weberi Osmólska, 1968 | Cph | Upper Pennsylvanian, Kasimovian | Russia, Vaigach Island |
| B. (A.) jesenicianus Hahn et Hahn in Hahn, Hahn et Ramovš, 1977 | Cc, Cph, Py | Upper Pennsylvanian, Gzhelian | Slovenia, Southern Karavanke |
| B. (A.) sp. Zhang, 1983 | Py | Pennsylvanian (?) | China, Xinjiang Uyghur Autonomous Region |
| B. (A.) chamberlaini (Kobayashi et Hamada, 1984a) | Py | Cisuralian, L. Wolfcampian (~Asselian), Phelan Creek Formation | USA, Alaska, vicinity of the Gulkan glacier |
| B. (A.) macgrathensis Hahn et Hahn, 1985 | Cph, Py | Upper Pennsylvanian – Cisuralian (Asselian) | USA, Alaska, Cheeneetnuk River Basin |
| B. (A.) edwardsi Owens, 1986 | Cph, Py | Lower Pennsylvanian, Kinderscoutian (~lower part of the Bashkirian) | England, West Yorkshire |
| B. (A.) spinicauda Gandl, 1987 | Cph, Py | Pennsylvanian, Namurian B – Westphalian (~Bashkirian-Moscovian) | Spain, Cantabrian Mountains, Palencia |
| B. (A.) acuticeps Gandl, 1987 | Cph, Py | Lower Pennsylvanian, Namurian C (~upper part of the Bashkirian) | Spain, Cantabrian Mountains, Leon |
| B. (A.) asiaticus Hahn, Hahn et Yuan, 1989 | Cph, Py | Lower Pennsylvanian, Dala Formation (~upper part of the Bashkirian) | China, Guangxi Zhuang Autonomous Region, Nandan County |
| B. (A.) junggarensis Wu et Feng, 1991 | Py | Upper Pennsylvanian, Shiqiantan Formation | China, Xinjiang Uygur Autonomous Region, Dzungaria |
| B. (A.) kalodermatus Hahn et Hahn, 1992 | Cph, Py | Lower Pennsylvanian, M. Morrowan – M. Atokan (~Bashkirian) | USA, Alaska, Alexander Archipelago |
| B. (A.) phalanx Gandl, 2011 | Cph, Py | Middle Pennsylvanian, Westphalian D (~upper part of the Moscovian) | Spain, Cantabrian Mountains, Palencia |
TABLE 2. Known species of Paraphillipsia Toumansky, 1930.
| Species | Part | Stratigraphy | Geography |
| P. ? middlemissi (Diener, 1897) | Py | Lopingian, Wuchiapingian (?), Chitichun Limestone | China, Tibet, Zanda County |
| P. ? sp. (Mansuy, 1912) | Py | Guadalupian (?) | Laos, Ban Na Hai |
| P. tschernyschewi (Netschaew in Weber, 1932) | Cc, Cph | Cisuralian, Safetdaron Formation | Tajikistan, Darvaz, Safed-Daron village, Tangi-Gor Gorge |
| P. pahara Weller, 1935 | Cr, Th, Py | Cisuralian (?), "reddish-gray coralline limestone" | China, Tibet, Eastern Karakoram, Chang Chenmo River valley |
| P. karpinskyi Tumanskaya, 1935 | Cc | Guadalupian, Roadian | Crimea, the Dzhien-Sofu Block on the Salgir River |
| P. baltensis Tumanskaya, 1935 | Py | Guadalupian, Roadian | Crimea, Kichkhi-Burnu Block on the Martha River |
| P. taurica Tumanskaya, 1935 | Cr, Lg, Py | Guadalupian, Roadian | Crimea, Kichkhi-Burnu Block on the Martha River |
| P. vnweberi Tumanskaya, 1935 | Cph, Py | Guadalupian, Roadian | Crimea, Kichkhi-Burnu Block on the Martha River |
| P.? sp. Weber, 1944 | Py | Guadalupian, Capitanian, Chandalazian horizon | Far East, Partizanskaya River, Sen’kina Shapka Mountain |
| P.? sp., aff. P. ? taurica Tumanskaya, 1935 (Hahn et Hahn in Hahn, Hahn et Ramovš, 1970 | Py | Cisuralian, Artinskian, Trogkofel Limestone | Slovenia, Karavanke, Dovžan Gorge |
| P.? sp. Hahn et Hahn, 1970 | Py | Cisuralian, Artinskian, Trogkofel limestone | Slovenia, Karavanke, Dovžan Gorge |
| P. levigata Kobayashi et Hamada, 1980 | Cr, Lg, Py | Guadalupian, Capitanian, Shimoyama Limestone, Yabeina zone | Japan, Shikoku, Sakawa |
| P. ? sinensis Zhou, 1987 | Cr, Th, Py | Cisuralian, Artinskian, Maping Formation | China, Guangxi Zhuang Autonomous Region, Bunuo |
| P.? carnica Schraut, 2019 | Py | Cisuralian, Artinskian, upper part of Zottachkopf Formation | Austria, Carnic Alps, surroundings of the Trogkofel and Troghöhe Mountains |
| P. urushtensis sp. nov. | Cph, Cr, Py | Lopingian, Changhsingian, Belalabino Group | North Caucasus, Malaya Laba River Basin |
TABLE 3. Dimensions (in mm) of the cephala and cranidia of Paraphillipsia urushtensis sp. nov. LC – length of the cephalon, WC – width of the cephalon, LG – length of the glabella, LO – length of the occipital ring, WGA – width of the glabella at the anterior end, WGP – width of the glabella at the posterior end, LL – length of the L1 lobes, LE – length of the eye.
| Specimen | LC | WC | LG | LO | WGA | WGP | LL | LE |
| CNIGRmuseum, No. 62/5217 | 7.7 | 8.8 | 6.4 | 4.1 | 5.8 | 5.5 | 2.8 | 2 |
| CNIGRmuseum, No. 61/5217 | - | - | 10.4 | 6.7 | 8 | 7 | 3.5 | - |
| CNIGRmuseum, No. 65/5217 | - | - | 8.9 | ~5 | 8 | 6.9 | 3.1 | - |
| CNIGRmuseum, No. 66/5217 | - | - | 6.4 | 4 | 5.1 | 4.4 | 2 | - |
TABLE 4. Dimensions (in mm) of pygidia of Paraphillipsia urushtensis sp. nov. LP – length of the pygidium, WP – width of the pygidium, WA – width of the axis at the anterior end, L/W – ratio of the length of the pygidium to its width, W/WA – ratio of the width of the pygidium to the width of the axis at the anterior end, DAB – distance from the end of the axis to edges of the border furrow of the pygidium.
| Specimen | LP | WP | WA | L/W | W/WA | DAB |
| CNIGRmuseum, No. 63/5217 | 6.7 | 7.5 | 3.8 | 0.9 | ~2 | 1 |
| CNIGRmuseum, No. 64/5217 | 7.5 | 9.6 | 5 | 0.8 | ~1.9 | 0.8 |
| CNIGRmuseum, No. 67/5217 | 6.9 | 7.3 | 3.8 | 0.9 | ~1.9 | 0.6 |
| CNIGRmuseum, No. 68/5217 | ~4 | ~4.4 | 2.3 | 0.9 | ~1.9 | 0.8 |
| CNIGRmuseum, No. 69/5217 | 5.6 | 6.7 | 3.4 | 0.8 | ~2 | >0.5 |
| Average value | ~6 | ~7 | ~3.7 | ~0.8 | ~1.9 | 0.8 |
TABLE 5. Type series of Paraphillipsia urushtensis sp. nov.
| Number of specimens | Part | Locality | Author of find, year | Photos |
| CNIGRmuseum, No. 61/5217 | Cr | Urushten tract (=paraje), outcrop No. 309 | V.N. Robinson, 1925 | Weber, 1944, pl. I, fig. 18; Hahn and Hahn, 2008, Abb. 333; Mychko, 2016, pl. III, fig. 6; herein – Figure 5 A, B |
| CNIGRmuseum, No. 62/5217 | Cph | 3 km from the estuary of the Urushten River, outcrop No. 265 | V.N. Robinson | Weber, 1944, pl. I, fig. 22; Hahn and Hahn, 2008, Abb. 332; Mychko, 2016, pl. III, fig. 3; herein – Figure 5 C |
| CNIGRmuseum, No. 63/5217 | Py | 3,5 km from the estuary of the Urushten River, outcrop No. 27 | B.K. Likharev, 1927 | Weber, 1944, pl. I, fig. 19; Hahn and Hahn, 2008, Abb. 334 a, b; Mychko, 2016, pl. III, fig. 5; herein – Figure 5 H |
| CNIGRmuseum, No. 64/5217 | Py | 3 km from the estuary of the Urushten River, outcrop No. 263 | V.N. Robinson | Weber, 1944, pl. I, fig. 20; Hahn and Hahn, 2008, Abb. 335; Mychko, 2016, pl. III, fig. 4; herein – Figure 5 F |
| CNIGRmuseum, No. 65/5217 | Cr | 3 km from the estuary of the Urushten River, outcrop No. 263 | V.N. Robinson | herein – Figure 5 D |
| CNIGRmuseum, No. 66/5217 | Cr | 3,5 km from the estuary of the Urushten River, outcrop No. 27 | B.K. Likharev, 1927 | herein – Figure 5 E |
| CNIGRmuseum, No. 67/5217 | Py | 3,5 km from the estuary of the Urushten River, outcrop No. 27 | B.K. Likharev, 1927 | herein – Figure 5 I |
| CNIGRmuseum, No. 68/5217 | Py | 3,5 km from the estuary of the Urushten River, outcrop No. 27 | B.K. Likharev, 1927 | herein – Figure 5 G |
| CNIGRmuseum, No. 69/5217 | Py | 2,25 from the estuary of the stream in Nikitinskaya Ravine | B.K. Likharev, 1927 | herein – Figure 5 J |
TABLE 6. Known species of Kathwaia Grant, 1966 adopted in this work.
| Species | Part | Stratigraphy | Geography |
| K. girtyi (Tumanskaya, 1935) | Th, Py | Guadalupian, Roadian | Crimea, Kichkhi-Burnu Block on the Martha River |
| K. (?) sinensis (Grabau, 1936) | Cc | Upper Pennsylvanian or Cisuralian, Maping Formation | China, Guangxi Zhuang Autonomous Region |
| K. caucasica (Weber, 1944) (including K. capitorosa Grant, 1966) | Cc, Cph, Cr, Gl, Py | Lopingian, Wuchiapingian –Changhsingian | Krasnodar Krai, Malaya Laba River Basin and Pakistan (Salt Range, Kathwai-Kushab) |
| K. ? kashmirensis (Sarkar 1968) | Py | Cisuralian (?) | India, Kashmir, vicinity of the Aishmuquam |
| K. ? sp. König et Kuss, 1980 | Cc | Cisuralian, Talea Ori Group | Greece, Crete, Bali village |
TABLE 7. Dimensions (in mm) of cephala, cranidia and glabellae of Kathwaia caucasica (Weber, 1944). LC – length of the cephalon, WC – width of the cephalon, LG – length of the glabella, LO – length of the occipital ring, WGA – width of the glabella at the anterior part, WGP – width of the glabella at the posterior part, LL – length of the L1 lobes, LE – length of the eye.
| Specimen | LC | WC | LG | LO | WGA | WGP | LL | LE |
| CNIGRmuseum, No. 53/5217 | ~12 | 16.9 | 10.2 | ~5.8 | 9.2 | 3.5 | 4 | 2.8 |
| USNM PAL 145320 | 8.5 | 11.7 | 6.7 | 5.6 | 6.2 | 2.6 | 2.7 | 1.5 |
| CNIGRmuseum, No. 54/5217 | - | - | 13.5 | ~8 | 10.9 | 4.7 | 4.3 | - |
| CNIGRmuseum, No. 55/5217 | - | - | - | - | 12.6 | - | - | - |
| CNIGRmuseum, No. 56/5217 | - | - | 9.2 | - | 7.4 | 2.6 | - | - |
| CNIGRmuseum, No. 57/5217 | - | - | 11.5 | - | 10 | 4.6 | - | - |
| CNIGRmuseum, No. 58/5217 | - | - | ~9 | - | 7 | ~3.5 | - | - |
| CNIGRmuseum, No. 59/5217 | - | - | 8 | - | 6.3 | 3 | - | - |
| Average value | 10.3 | 14.3 | 9.73 | 6.5 | 8.7 | 3.5 | 3.7 | 2.2 |
TABLE 8. Dimensions (in mm) of pygidia of Kathwaia caucasica (Weber, 1944). LP – length of the pygidium, WP – width of the pygidium, WA – width of the axis at the anterior end, L/W – ratio of the length of the pygidium to its width, W/WA – ratio of the width of the pygidium to the width of the axis at the anterior end, DAB – distance from the end of the axis to edges of the border furrow of the pygidium.
| Specimen | LP | WP | WA | L/W | W/WA | DAB |
| CNIGRmuseum, No. 87/5217 | 12 | ~17 | 8 | 0.7 | 2.1 | 2 |
| CNIGRmuseum, No. 88/5217 | 8 | 11.3 | 4.3 | 0.7 | 2.6 | 1.4 |
| USNM PAL 145320 | 7.9 | 11.3 | 4.5 | 0.7 | 2.5 | 1 |
| Average value | 9.3 | 13.2 | 5.6 | 0.7 | 2.4 | 1.5 |
TABLE 9. Hypodygm (type series and hypotype) of Kathwaia caucasica (Weber, 1944).
| Number | Part | Locality | Author of finding, year | Photo |
| CNIGRmuseum, No. 87/5217 | Py | Raskol-Skala Mountain, near Khamyshki Village, western part of the block, outcrop No. 42 | B.K. Likharev, 1927 | Weber, 1944, pl. II, fig. 16; Mychko, 2016, pl. II, fig. 17; herein – Figure 7 J |
| CNIGRmuseum, No. 88/5217 | Py | 3 km from Urushten tract (=paraje), outcrop No. 264 | V.N. Robinson | Weber, 1944, pl. II, fig. 17; Hahn and Hahn, 2015, abb. 110; Mychko, 2016, pl. II, fig. 18; herein – Figure 7 K |
| CNIGRmuseum, No. 53/5217 | Cph | Gefo Mountain, blocks along the Tegen’ River, outcrop No. 30 | B.K. Likharev, 1927 | Weber, 1944, pl. I, fig. 15; Hahn and Hahn, 2015, abb. 109; Mychko, 2016, pl. I, fig. 20; herein – Figure 7 A, B |
| CNIGRmuseum, No. 54/5217 | Cr | Raskol-Skala Mountain, near Khamyshki Village, western part of the block, outcrop No. 42 | B.K. Likharev, 1927 | Weber, 1944, pl. I, fig. 16; Mychko, 2016, pl. I, fig. 19; herein – Figure 7 C, D |
| CNIGRmuseum, No. 55/5217 | Gl | - // - | - // - | herein – Figure 7 E |
| CNIGRmuseum, No. 56/5217 | Gl | - // - | - // - | herein – Figure 7 H |
| CNIGRmuseum, No. 57/5217 | Gl | - // - | - // - | herein – Figure 7 G |
| CNIGRmuseum, No. 58/5217 | Gl | - // - | - // - | herein – Figure 7 F |
| CNIGRmuseum, No. 59/5217 | Gl | - // - | - // - | herein – Figure 7 I |
| USNM PAL 145320 | Cc | Pakistan, Salt Range, near the Kathwai Village | A.N. Fatmi, 1963-1964 | Grant, 1966, pl. 13, fig. 1a-d; Owens, 1983, pl. 2, fig. 1-4; Hahn and Brauckmann, 1988, Taf. 2, fig. 20-21; Owens, 2003, text-fig. 3A-B; Hahn and Hahn, 2015, Abb. 108-109; herein – Figure 8 A-D |
TABLE 10. Known species of Pseudophillipsia accepted in this work (those described exclusively by pygidia are conditionally classified as Pseudophillipsia).
| Species | Part | Stratigraphy | Geography |
| Ps. sumatrensis (Roemer, 1880) | Cc, Hy | Guadalupian, Wordian | Indonesia, the western coast of Sumatra |
| Ps. obtusicauda (Kayser, 1883) | Cr, Py | Lopingian | Southern China |
| Ps. (?) oehlerti (Gemmellaro, 1892) | Py | Guadalupian, Roadian | Italy, Sicily, Sosio River Valley |
| Ps. elegans Gemmellaro, 1892 | Cph, Th, Py | Guadalupian, Roadian | Italy, Sicily, Sosio River Valley |
| Ps. (?) sosiensis (Gemmellaro, 1892) | Py | Guadalupian, Roadian | Italy, Sicily, Sosio River Valley |
| Ps. (?) acuminata Mansuy, 1912 | Py | Guadalupian, graywacke with Lyttonia cf. tenuis | Laos, Luang Prabang (Ban-Pak-Luang and Xieng Men) |
| Ps. gemmellaroi Canavari in Greco, 1935 | Cc | Guadalupian, Roadian | Italy, Sicily, Sosio River Valley |
| Ps. borissiaki Tumanskaya, 1935 | Cc | Guadalupian, Roadian | Crimea, Dzhien-Sofu (=Totai-Koi) Block on the Salgir River |
| Ps. (?) crimensis Tumanskaya, 1935 | Py | Guadalupian, Roadian | Crimea, Kichkhi-Burnu Block on the Marta River |
| Ps. (?) ibrischensis Tumanskaya, 1935 | Py | Guadalupian, Roadian | Crimea, Kichkhi-Burnu Block on the Marta River |
| Ps. (?) martensis Tumanskaya, 1935 | Py | Guadalupian, Roadian | Crimea, Kichkhi-Burnu Block on the Marta River |
| Ps. (?) mustafensis Tumanskaya, 1935 | Py | Guadalupian, Roadian | Crimea, Dzhien-Sofu (=Totai-Koi) Block on the Salgir River |
| Ps. (?) sarabensis Tumanskaya, 1935 | Py | Guadalupian, Roadian | Crimea, Kichkhi-Burnu block on the Marta River |
| Ps. (?) timorensis (Gheyselinck, 1937) | Py | Guadalupian | Indonesia, East Nusa Tenggara Province, Basleo faunas |
| Ps. solida Weber, 1944 | Cph, Cr | Lopingian, Changhsingian | Russia, Krasnodar Krai and Slovenia |
| Ps. (?) cf. mustafensis Tumanskaya, 1935 (in Weber, 1944) | Py | Lopingian, Changhsingian | Russia, Krasnodar Krai |
| Ps. (?) caucasica Weber, 1944 | Py | Lopingian, Changhsingian | Russia, Krasnodar Krai |
| Ps. (?) armenica Weber, 1944 | Py | Guadalupian | Armenia and Iran |
| Ps. (?) hungarica (Schréter, 1948) | Cr, Lg, Py | Lopingian, Nagyvisnyó Formation | Hungary, Bükk Mountains |
| Ps. aff. sumatrensis (Roemer, 1880) (in Hahn et al., 1970) | Cph, Py | Lopingian, Changhsingian, Bellerophon Formation | Slovenia, Vrzdenec |
| Ps. (?) cf. hungarica (Schréter, 1948) (in Hahn et al., 1970) | Py | Lopingian, Changhsingian, Bellerophon Formation | Slovenia, Žažar |
| Ps. azzouzi Termier et Termier 1974 | Cc | Guadalupian, Capitanian | Tunisie, Djebel Tebaga |
| Ps. anshunensis Qian, 1977 | Cph, Py | Lopingian, Wuchiapingian | China, Guizhou, Jiaozishan Section |
| Ps. (?) subcircularis Qian, 1977 | Py | Lopingian, Wuchiapingian | China, Guizhou, Jiaozishan Section |
| Ps. qinglongensis Qian, 1977 | Cph+Tx, Fr | Lopingian, Changhsingian, Dalong Formation | China, Guizhou, Zhongying Section |
| Ps. (?) huishuiensis Yin, 1978 | Py | Cisuralian, Kungurian | China, Chengfanguan, Huishui County |
| Ps. tongluensis Ju in Zhang, 1982 | Cc, Py | Guadalupian, Wordian-Capitanian, Dingjiashan Formation | China, Tonglu |
| Ps. (?) wuweiensis Zhang, 1982 | Py | Guadalupian, Qixia Formation | China, Anhui, Wuwei County |
| Ps. shanggaoensis Zhang, 1982 | Cr | Lopingian, Wuchiapingian | China, Jiangxi, Shanggao County and Gao'an County, Loping Formation |
| Ps. yunanxiensis Liu, 1982 | Cc | Cisuralian (?) | China, Sanzhi Count，Yunanxi |
| Ps. akasakensis Kobayashi et Hamada, 1984b | Cph+Py | Guadalupian, Roadian-Capitanian Neoschwagerina Zone (?) – Yabeina Zone | Japan, Gifu Prefecture, Akasaka Limestone |
| Ps. ozawai Kobayashi et Hamada, 1984b | Cr, Py | Guadalupian, Capitanian, Yabeina Zone | Japan, Gifu Prefecture, Akasaka Limestone |
| Ps. aff. ozawai Kobayashi et Hamada, 1984b (in Kobayashi abd Sakagami, 1989) | Py | Lopingian, Changhsingian, Huai Thak Formation | Thailand, Lampang Province, Doi Pha Phlung |
| Ps. kiriuensis Kobayashi et Hamada, 1984b | Cr, Py | Guadalupian, Roadian-Capitanian, Nabeyama Formation Parafusulina Zone | Japan, Gunma and Gifu Prefecture |
| Ps. (?) subtrigonalis Kobayashi et Hamada, 1984a | Py | Guadalupian, Roadian-Capitanian, Nabeyama Formation | Japan, Tochigi Prefecture, Kuzuu |
| Ps. (?) kuzuensis Kobayashi et Hamada, 1984a | Py | Guadalupian Yamasuge imestone, Nabeyama Formation | Japan, Tochigi Prefecture, Kuzuu |
| Ps. hanaokensis Kobayashi et Hamada, 1984b | Cr, Py | Lopingian, Ichihashi Formation | Japan, Gifu Prefecture, Akasaka. Akasaka Limestone. |
| Ps. (?) binodosa Kobayashi et Hamada, 1984a | Py | Guadalupian, Wordian, Shigejizawa Formation | Japan, Anabuchi, Kesennuma City, Miyagi Prefecture, Myogo-sawa |
| Ps. sasakii Kobayashi et Hamada, 1984a | Cr, Py | Guadalupian, Wordian, Shigejizawa Formation | Japan, Anabuchi, Kesennuma City, Miyagi Prefecture, Myogo-sawa |
| Ps. (?) simplex Kobayashi et Hamada, 1984a | Py | Guadalupian, Wordian, Shigejizawa Formation | Japan,Anabuchi, Kesennuma City, Miyagi Prefecture, Myogo-sawa |
| Ps. spatulifera Kobayashi et Hamada, 1984a | Cc, Cr, Py | Guadalupian, Capitanian | Japan,Kanokura Formation (various localities) |
| Ps. catena Kobayashi et Hamada, 1984b | Cr, Py, Fr | Guadalupian, Roadian-Capitanian, Neoschwagerina Zone (?) – Yabeina Zone | Japan, Gifu Prefecture, Akasaka Limestone |
| Ps. (?) perturbata Hahn, Hahn et Ramovš 1990 | Py | Guadalupian, Roadian | Crimea, Kichkhi-Burnu Block on the Marta River |
| Ps. pradilla Gandl, 2011 | Cr, Py | Middle Pennsylvanian, Moscovian (Westphalian D) | Spain, Palencia, Blatt Cervera de Pisuerga, |
| Ps. (?) aff. caucasica Weber, 1944 (in Lerosey-Aubril, 2012) | Py | Lopingian, Nesen Formation | Iran, Alborz range, Yush |
| Ps. (?) parvizii Lerosey-Aubril, 2012 | Py | Lopingian, Wuchiapingian, Dalan Formation | Iran, Dena Mountain, northwest of Yasouj |
| Ps. darvazica Mychko, 2016 nom. nud. | Cph | Cisuralian, Artinskian, Safetdaron Formation | Tajikistan, Darvaz, Obihingou River |
TABLE 11. Dimensions (in mm) of cephala, cranidia and glabellae of Pseudophillipsia solida Weber, 1944. LC – length of the cephalon, WC – width of the cephalon, LG – length of the glabella, LO – length of the occipital ring, WGA – width of the glabella at the anterior part, WGP – width of the glabella at the posterior part, LL – length of the L1 lobes, LE – length of the eye.
| Specimen | LC | WC | LG | LO | WGA | WGP | LL | LE |
| CNIGRmuseum, No. 79/5217 | 9 | 12 | 5.8 | ~1.5 | 5.5 | 2 | 1.3 | 2.4 |
| CNIGRmuseum, No. 80/5217 | - | - | 9.5 | >2 | 7.5 | 3.4 | - | - |
| CNIGRmuseum, No. 81/5217 | - | - | ~5 | - | 4.5 | - | - | - |
| GPIUL, No. 3853 | - | - | 7.4 | >3 | 6 | 2.5 | - | - |
| Average value | ~9 | ~12 | 7.6 | 1.5 | 5.8 | 3 | ~1.3 | 2.4 |
TABLE 12. Known specimens of Pseudophillipsia (?) solida Weber, 1944; *localities at the North Caucasus (Malaya Laba River Basin).
| Number of specimens | Part | Locality | Author of find, year | Photos |
| CNIGRmuseum, No. 79/5217 | Cph | *3,05 km from the estuary of the Urushten River, outcrop No. 127 | B.K. Likharev, 1927 | Weber, 1944, pl. II, fig. 8; Mychko 2016, pl. VI, fig. 6; herein – Figure 10A, B |
| CNIGRmuseum, No. 80/5217 | Cr | *3,05 km from the estuary of the Urushten River, outcrop No. 127 | B.K. Likharev, 1927 | Weber, 1944, pl. II, fig. 9; Mychko 2016, pl. VI, fig. 7; herein – Figure 10C |
| CNIGRmuseum, No. 81/5217 | Cr | *Gefo Mount, outcrop No. 30 33 c. | B.K. Likharev, 1927 | herein – Figure 10D |
| GPIUL, No. 3853 | Cr | Slovenia, vicinity of the Vrzdenec Village | A. Ramovš, before 1970 | Hahn, Hahn et Ramovš, 1970, pl. 1, fig. 4; herein – Figure 10D |
TABLE 13. Dimensions (in mm) of pygidia of Pseudophillipsia (?) caucasica Weber, 1944. LP – length of the pygidium, WP – width of the pygidium, WA – width of the axis at the anterior part, L/W – ratio of the length of the pygidium to its width, W/WA – ratio of the width of the pygidium to the width of the axis at the anterior part, DAB – distance from the end of the axis to edges of the border furrow of the pygidium. *the deformed (compressed from the sides) specimen.
| Specimen | LP | WP | WA | L/W | W/WA | DAB |
| CNIGRmuseum, No. 71/5217 | 8.1 | >8 | ~3.3 | ~1 | ~2.4 | 0.4 |
| CNIGRmuseum, No. 72/5217 | 18.7 | ? | ? | ? | ? | ? |
| CNIGRmuseum, No. 73/5217 | 14.7 | ~14 | 4.8 | ~1 | ~2.9 | 1.3 |
| CNIGRmuseum, No. 74/5217* | ~9.2 | ~7 | ~2 | <1.3 | <3.5 | ~1 |
| Average value | ~12.7 | ~9.7 | 3.4 | ~1 | ~2.9 | ~0.9 |
TABLE 14. Known specimens of Pseudophillipsia (?) caucasica Weber, 1944; *incomplete specimen.
| Number of specimens | Part | Locality | Author of find, year | Photos |
| CNIGRmuseum, No. 71/5217 | Py* | Gefo Mount, blocks along the Tegen’ River | V.N. Robinson, 1925 | Weber, 1944, pl. II, fig. 4; Mychko 2016, pl. VI, fig. 2; herein – Figure 10 E |
| CNIGRmuseum, No. 72/5217 | Py | 2,25 from the estuary of the stream in Nikitinskaya Ravine | B.K. Likharev, 1927 | herein – Figure 10 F |
| CNIGRmuseum, No. 73/5217 | Py | Urushten tract (=paraje), outcrop No. 309 | V.N. Robinson, 1925 | Weber, 1944, pl. II, fig. 2; non (!) Lerosey-Aubril, 2012, p. 10, fig. 4 A; Mychko 2016, pl. VI, fig. 1; herein – Figure 10 G |
| CNIGRmuseum, No. 74/5217 | Py | 2,25 from the estuary of the stream in Nikitinskaya Ravine | B.K. Likharev, 1927 | herein – Figure 10 H |
TABLE 15. Dimensions (in mm) of pygidium of Pseudophillipsia (?) cf. mustafensis Tumanskaya, 1935. LP – length of the pygidium, WP – width of the pygidium, WA – width of the axis at the anterior end, L/W – ratio of the length of the pygidium to its width, W/WA – ratio of the width of the pygidium to the width of the axis at the anterior end, DAB – distance from the end of the axis to edges of the border furrow of the pygidium.
| Specimen | LP | WP | WA | L/W | W/WA | DAB |
| CNIGRmuseum, No. 77/5217 | 7.5 | ~8.6 | 2.7 | ~0.9 | ~3 | 0.4 |
TABLE 16. Known specimens of Pseudophillipsia (?) cf. mustafensis Tumanskaya, 1935; *incomplete specimen, fragment of a lateral lobe.
| Number of specimens | Part | Locality | Author of find, year | Photos |
| CNIGRmuseum, No. 77/5217 | Py | Gefo Mount, outcrop No. 30 | B.K. Likharev, 1927 | Weber, 1944, pl. II, fig. 3; herein – Figure 10 I |
| CNIGRmuseum*, No. 78/5217 | Py | Gefo Mount, outcrop No. 30 | B.K. Likharev, 1927 | herein – Figure 10 J |
TABLE 17. Distribution of Lopingian trilobites in the World. Lop – undivided Lopingian deposits, Wu – Wuchiapingian, Ch – Changhsingian.
| Locality | Formation | Stage | Species / forms | References |
| Vrzdenec (Slovenia) | Bellerophon Fm. (formed Žažar Fm.) | Ch. | Pseudophillipsia n. sp., aff. sumatrensis (Roemer, 1880) Pseudophillipsia solida Weber, 1944 |
Hahn et al., 1970; Hahn, Hahn et Brauckman, 2001 |
| Žažar (Slovenia) | Bellerophon Fm. (former Žažar Fm.) | Ch. | Pseudophillipsia (?) cf. hungarica (Schréter, 1948) | Hahn et al., 1970; Hahn, Hahn, Brauckman, 2001 |
| Bükk Mountains (Hungary) | Nagyvisnyó Fm. | Ch. | Pseudophillipsia (?) hungarica (Schréter, 1948) | Schréter, 1948; Detre, 1991 |
| Crimea, blocks on the river Alma | - | Lop.? | Pseudophillipsia (?) sp. ind. № 1 Pseudophillipsia (?) sp. ind. № 2 Pseudophillipsia sp. ind. № 1 |
Tumanskaya, 1935 |
| Far East, Nakhodka | Lyudyanzian | Wu. | Paraphillipsia sp. Neogriffithides (?) sp. |
Herein |
| North Caucasus | Nikitino and Urushten Fm’s. | Ch. | Brachymetopus (Acutimetopus) caucasicus Licharew in Weber, 1944 Kathwaia caucasica (Weber, 1944) Paraphillipsia urushtensis sp. nov. Pseudophillipsia (?) caucasica Weber, 1944 Pseudophillipsia mustafensis Tumanskaya, 1935 Pseudophillipsia solida Weber, 1944 |
Weber, 1944; this research |
| Alborz range, Yush (Iran) | Nesen Fm. | Wu.-Ch. | Acropyge weggeni Hahn et Hahn, 1981 Pseudophillipsia aff. caucasica Weber, 1944 |
Hahn and Hahn, 1981; Hahn, Hahn et Brauckman, 2001; Lerosey-Aubril, 2012 |
| Dena Mountain, northwest of Yasouj (Iran) | Dalan Fm. | Wu. | Pseudophillipsia (?) parvizii Lerosey-Aubril, 2012 | Lerosey-Aubril, 2012 |
| Salt Range, Zaluch Nala-Kala Wahan (Pakistan) | Wargal Fm. | Wu. | Ditomopyge (Carniphillipsia) fatmii Grant, 1966 | Grant, 1966; Hahn, Hahn et Brauckman, 2001 |
| Salt Range, Kathwai-Kushab (Pakistan) | Wargal Fm. | Wu. | Kathwaia caucasica (Weber, 1944) (= Kathwaia capitorosa Grant, 1966) | Grant, 1966; Hahn, Hahn et Brauckman, 2001; this research |
| Tibet, Zanda County (China) | Chitichun Limestone | Wu. (?) | Cheiropyge himalayensis Diener, 1897 Paraphillipsia (?) middlemissi Diener, 1897 |
Diener, 1897; Hahn, Hahn et Brauckman, 2001 |
| Tibet, Shuanghu (China) | Raggyorcaka Fm. | Ch. (?) | Ditomopyge (Carniphillipsia) raggyorcakaensis (Qian 1981) | Qian, 1981; Hahn, Hahn et Brauckman, 2001 |
| Chongqing, Beifengjing Section (China) | Changxing Fm. | Ch. | Pseudophillipsia obtusicauda (Kayser, 1883) Ditomopyge (Carniphillipsia) chongqingensis (Lu, 1974) Pseudophillipsia sp. |
Shen and He, 1991 |
| Chongqing, Tudiya buildup (China) | Changxing Fm. | Ch. | Ditomopyge (Carniphillipsia) cf. chongqingensis (Lu, 1974) | Reinhardt, 1988 |
| Chongqing, Daijiagou Beipei, Yanjingxi section (China) | Changxing Fm. | Ch. | Pseudophillipsia (?) sp. | Shen et al., 1995; Yang et al., 1987 |
| Chongqing, Liziya section (China) | Longtan Fm. | Wu. | Pseudophillipsia (?) sp. | Zeng et al., 1995 |
| Chongqing, Huaying Section (China) | Changxing Fm. | Ch. | Pseudophillipsia obtusicauda (Kayser, 1883) Ditomopyge (Carniphillipsia) chongqingensis (Lu, 1974) |
Yang et al., 1987 |
| Chongqing, Wenxing (China) | Longtan Formation | Wu. | Ditomopyge (Carniphillipsia) lui (Kobayashi et Hamada, 1984a) Ditomopyge (Carniphillipsia) chongqingensis (Lu, 1974) |
Lu, 1974; Kobayashi and Hamada, 1984a |
| Chongqing, Zhongliangshan (China) | Changxing Fm. | Ch. | Ditomopyge (Carniphillipsia) cf. chongqingensis (Lu, 1974) | Shi et al., 2016 |
| Guizhou, Jiaozishan Section (China) | Longtan Fm. | Wu. | Pseudophillipsia anshunensis Qian, 1977 Pseudophillipsia (?) subcircularis Qian, 1977 |
Qian, 1977; Kobayashi and Hamada, 1984a |
| Guizhou, Yanbeihou (China) | Longtan Fm. | Wu. | Pseudophillipsia (?) sp. | Wang et al., 2011 |
| Guizhou, Xinmin section (China) | Dalong Fm. | Ch. | Ditomopyge (Carniphillipsia) cf. chongqingensis (Lu, 1974) | Feng et al., 2011 |
| Guizhou, Tianshengqiao (China) | Dalong Fm. | Ch. | Acropyge brevica Yin, 1978 | Yin, 1978 |
| Guizhou, Wenjiangsi Section (China) | Changxing Fm. | Ch. | Pseudophillipsia obtusicauda (Kayser, 1883) Ditomopyge (Carniphillipsia) chongqingensis (Lu, 1974) |
Shen and He, 1994 |
| Guizhou, Zhongying Section (China) | Dalong Fm. | Ch. | Pseudophillipsia qinglongensis Qian, 1977 Ditomopyge (Carniphillipsia) pyriformis (Qian, 1977) Acropyge multisegmenta Qian, 1977 |
Qian, 1977 |
| Guangxi, Heshan Section and Paoshui Section (China) | Heshan Fm. | Wu. | Ditomopyge (Carniphillipsia) heshanensis (Qian, 1977) Pseudophillipsia obtusicauda (Kayser, 1883) |
Qian, 1977; Yang et al., 1987 |
| Guangdong, Qujiang County (China) | Changxing Fm. | Ch. | Ditomopyge (Carniphillipsia) lui (Kobayashi et Hamada, 1984a) | Zhou, 1977; Kobayashi and Hamada, 1984a |
| Jiangxi, Shanggao County and Gao'an County (China) | Loping Formation, Laoshan Member | Wu. | Pseudophillipsia shanggaoensis Zhang, 1982 Cheiropyge (?) gaoanensis Zhang, 1982 |
Zhang, 1982 |
| Jiangxi, Shanggao County and Leping County (China) | ? | Lop. | Pseudophillipsia obtusicauda Kayser, 1883 | Kayser, 1883; Zhang, 1982 |
| West Timor, Kali-Mati (Indonesia) | Amarassi Beds | Wu. | Timorcranium parvulum (Beyrich, 1865) | Beyrich, 1865; Hahn et Brauckmann, 1975; Brauckmann and Gröning, 2013 |
| Gifu Prefecture, Akasaka (Japan) | Akasaka Limestone, Ichihashi Fm. | Lop. | Pseudophillipsia hanaokensis Kobayashi et Hamada, 1984b | Kobayashi and Hamada, 1984a; 1984b |
| Lampang Province, Doi Pha Phlung and Huai Mae Phlung (Thailand) | Huai Thak Fm. | Ch. | Pseudophillipsia aff. ozawai Kobayashi et Hamada, 1984a | Kobayashi and Sakagami, 1989; Ishibashi, 1998 |
| Mossburn, Oreti River (New Zealand) | Countess Fm., Stephens Subgroup | Ch. (?) | Triproetus sp. | Hyden et al., 1982 |
| Spitsbergen, Ahlstrandhalvøya peninsula | Kapp Starostin Fm., Hovtinden member | Lop. | Triproetus borealis Kobayashi, 1987 | Kobayashi, 1987 |
TABLE 18. Lopingian species of Ditomopyge (Carniphillipsia).
| Species | Images | Part | Axial rings | Pleural ribs |
| D. (C.) fatmii Grant, 1966 | Grant 1966, pl. 13, fig. 2,4; herein – Figure 12M.1, 12M.2 | Cc | 17 | 12 |
| D. (C.) chongqingensis (Lu, 1974) | Lu, 1974, pl. 166, figs. 27,28; Lerosey-Aubril, 2012, fig. 6 d, g, h, k, m, p; herein – Figure 12N.1, 12.N.2 | Cph, Py | 25 | 14-15 |
| D. (C.) heshanensis (Qian, 1977) | Qian, 1977, pl. 1, fig. 10 herein – Figure 12O | Cc | 21 | 13-14 |
| D. (C.) pyriformis (Qian, 1977) | Qian, 1977, pl. 1, fig. 10; herein – Figure 12R (only cranidium) | Cph | ? | ? |
| D. (C.) raggyorcakaensis (Qian, 1981) | Qian, 1981, pl. 1, figs. 5,6; herein – Figure 12Q (only cranidium) | Cr, Py | 20 | 12 |
| D. (C.) lui (Kobayashi et Hamada, 1984a) | Lu, 1974, pl. 166, figs. 25,26; herein – Figure 12P | Cr, Th+Py | 22-23 | 13-14 |
TABLE 19. Lopingian species of Pseudophillipsia.
| Species | Images | Part | Axial rings | Pleural ribs |
| Ps. anshunensis Qian, 1977 | Qian, 1977, pl. I, figs. 4,5; herein – Figure 12U.1, 12U.2 | Cph, Py | 22 | 12 (13-14) |
| Ps. qinglongensis Qian, 1977 | Qian, 1977, pl. I, figs. 1-3; herein – Figure 12Y (only cephalon) | Cph, Th, Py* | ? | ? |
| Ps. shanggaoensis Zhang, 1982 | Zhang, 1982, pl. 125, fig. 2; herein – Figure 12AA | Cr | ? | ? |
| Ps. obtusicauda (Kayser, 1883) | Zhang, 1982, pl. 125, fig. 1; herein – Figure 12AC.1, 12AC.2 | Cc | 20 | 12 |
| Ps. hanaokensis Kobayashi et Hamada, 1984b | Kobayashi and Hamada, 1984a, pl. X, figs. 1-5; pl. XI, figs. 3,4; Text-fig. 6-a; Kobayashi and Hamada, 1984b, Figure 5; herein – Figure 12Z.1, 12Z.2 | Cr, Py | 25-24 | 10-11 |
| Ps. n. sp., aff. sumatrensis (Roemer, 1880) | Hahn et al., 1970, Taf. 1, fig. 7, abb. 4; herein – Figure 9.24 | *Cph, *Py | ? | ~12(?) |
| Ps. solida Weber, 1944 | Weber, pl. II, fig. 8,9; herein – Figure 10 a-d, Figure 12V, 12W | Cph | ? | ? |
| Ps. (?) aff. ozawai Kobayashi et Hamada, 1984b | Kobayashi and Sakagami, 1989, fig. 1; herein – Figure 12AB | Py | 25-24 | 17 |
| Ps. (?) hungarica (Schréter, 1948) | Schréter, 1948, fig. 4; Detre, 1991, figs. 2,3; herein – Figure 12S | Py | 27 | 13 |
| Ps. (?) cf. hungarica (Schréter, 1948) | Hahn et al., 1970, fig. 3; herein – Figure 12T | Py | >18 | 12(13-14) |
| Ps. (?) parvizii Lerosey-Aubril, 2012 | Lerosey-Aubril, 2012, fig. 5; herein – Figure 12AF | Py | 21 | 13 |
| Ps. (?) subcircularis Qian, 1977 | Qian, 1977, pl. I, figs. 7,8; herein – Figure 12AD | Py | 23-24 | 12 |
| Ps. (?) caucasica Weber, 1944 | Weber, 1944, pl. II, fig. 4; herein – Figure 10 a-d, Figure 12AG | Py | 25 | 11 |
| Ps. (?) aff. caucasica Weber, 1944 | Lerosey-Aubril, 2012, figs. 4g-j, l, m; herein – Figure 12AH | Py | >17 (21-22) | 10 (?) |
| Ps. (?) cf. mustafensis Tumanskaya, 1935 | Weber, 1944, pl. II, fig. 3; herein – Figure 10 i-k, Figure 12AE | Py | 25 | 12 |
TABLE 20. Taxonomy of Lopingian trilobites adopted in this article. *primary identifications of unpublished trilobites from the Wuchiapingian of the Russian Far East.
| Brachymetopidae Prantl et Přibyl, 1950 | |||
| Brachymetopus McCoy, 1847 | |||
| Brachymetopus (Acutimetopus) Hahn et Hahn, 1985 | |||
| Brachymetopus (Acutimetopus) caucasicus Licharew in Weber, 1944 | |||
| Cheiropyge Diener, 1897 | |||
| Cheiropyge himalayensis Diener, 1897 | |||
| Cheiropyge (?) gaoanensis (Zhang, 1982) | |||
| Proetidae Hawle et Corda, 1847 | |||
| Triproetus Kobayashi and Hamada, 1979 | |||
| Triproetus borealis Kobayashi, 1987 | |||
| Triproetus sp. | |||
| Phillipsiidae Oehlert, 1886 | |||
| Cummingellinae Hahn et Hahn, 1967 | |||
| Paraphillipsia Toumansky, 1930 | |||
| Paraphillipsia urushtensis sp. nov. | |||
| Paraphillipsia (?) middlemissi Diener, 1897 | |||
| Paraphillipsia sp.* | |||
| Timorcranium Brauckmann et Gröning, 2013 | |||
| Timorcranium parvulum (Beyrich, 1865) | |||
| Bollandiinae Hahn et Brauckmann, 1988 | |||
| Kathwaia Grant, 1966 | |||
| Kathwaia caucasica (Weber, 1944) | |||
| Neogriffithides Toumansky, 1930 | |||
| Neogriffithides (?) sp.* | |||
| Ditomopyginae Hupé, 1953 | |||
| Acropyge Qian, 1977 | |||
| Acropyge brevica Yin, 1978 | |||
| Acropyge multisegmenta Qian, 1977 | |||
| Acropyge weggeni Hahn et Hahn, 1981 | |||
| Ditomopyge Newell, 1931 | |||
| Ditomopyge (Carniphillipsia) Hahn et Brauckmann, 1975 | |||
| Ditomopyge (Carniphillipsia) fatmii Grant, 1966 | |||
| Ditomopyge (Carniphillipsia) chongqingensis (Lu, 1974) | |||
| Ditomopyge (Carniphillipsia) heshanensis (Qian, 1977) | |||
| Ditomopyge (Carniphillipsia) pyriformis (Qian, 1977) | |||
| Ditomopyge (Carniphillipsia) raggyorcakaensis (Qian, 1981) | |||
| Ditomopyge (Carniphillipsia) lui (Kobayashi et Hamada, 1984a) | |||
| Pseudophillipsia Gemmellaro, 1892 | |||
| Pseudophillipsia anshunensis Qian, 1977 | |||
| Pseudophillipsia hanaokensis Kobayashi et Hamada, 1984b | |||
| Pseudophillipsia n. sp., aff. sumatrensis (Roemer, 1880) | |||
| Pseudophillipsia obtusicauda (Kayser, 1883) | |||
| Pseudophillipsia qinglongensis Qian, 1977 | |||
| Pseudophillipsia shanggaoensis Zhang, 1982 | |||
| Pseudophillipsia solida Weber, 1944 | |||
| Pseudophillipsia (?) aff. ozawai Kobayashi et Hamada, 1984b | |||
| Pseudophillipsia (?) caucasica Weber, 1944 | |||
| Pseudophillipsia (?) cf. mustafensis Tumanskaya, 1935 | |||
| Pseudophillipsia (?) aff. caucasica Weber, 1944 | |||
| Pseudophillipsia (?) parvizii Lerosey-Aubril, 2012 | |||
| Pseudophillipsia (?) hungarica (Schréter, 1948) | |||
| Pseudophillipsia (?) cf. hungarica (Schréter, 1948) | |||
| Pseudophillipsia (?) subcircularis Qian, 1977 | |||
| Pseudophillipsia (?) sp. | |||
