FIGURE 1. Geographic and geological setting and general stratigraphy of the Galve sub-basin in the Camarillas area (based on Aurell et al., 2016; Liesa et al., 2023), including the location of the Fuente Arnar site, type locality of Camarillasaurus cirugedae.
FIGURE 2. Reconstruction of Camarillasaurus cirugedae with preserved bones indicated. Note that some elements are from the right part of the skeleton (see text for details). Scale bar indicates 1 m.
FIGURE 3. Fused right surangular/articular of Camarillasaurus cirugedae, MPZ 2022/182b. A, lateral view (stereophotograph); B, posterior view; C, anterior view; D, dorsal view. Abbreviations: dp, dorsal process; f, foramen; g, groove; gl, concave facets of the mandibular glenoid; lds, laterodorsally facing shelf for the attachment of the m. aductor mandibulae externus superficialis; lsr, lateral surangular ridge; lt, lateral tubercle; r, ridge separating the mandibular glenoid cotyles; ri, ridge bordering the glenoid anterolaterally; rp, retroarticular process. Scale bar is 5 cm.
FIGURE 4. Teeth referred to Camarillasaurus cirugedae. A-C, MPG-KPC 43, tooth tip in ?labial (A), ?lingual (B) and ?distal views. D-F, MPZ 2022/182e, partial tooth in labial (D), lingual (E), mesial (F), apical (G) and basal (H) views. Abbreviations: dc, distal carina; pt, protuberance. Scale bars are 5 mm.
FIGURE 5. Remains of dorsal vertebrae of Camarillasaurus cirugedae. A, B, isolated anterior dorsal left prezygapophysis (MPG-KPC 40) in lateral (A) and anterior (B) views. C-E, posterior dorsal vertebral centrum and partial neural arch (MPG-KPC 9) in left lateral (C), anterior (D) and ventral (E) views. F, G, isolated left transverse process (MPG-KPC 14) in anterior (F) and ventral (G) views. Abbreviations: cprl, centroprezygapophyseal lamina; di, diapophysis; hyp, hypantrum; pap, parapophysis; pcdl, posterior centrodiapophyseal lamina; ppdl, paradiapophyseal lamina; prz, prezygapophysis. Scale bars are 5 cm.
FIGURE 6. Sacral vertebrae and possible posterior dorsal or sacral neural spine of Camarillasaurus cirugedae. A, B, probable partial primordial sacral centra one and two and a caudal sacral vertebral centrum (MPG-KPC 3 and 4) in left lateral (A) and ventral (B) views. Anterior is to the left. C, D, posterior dorsal or sacral neural spine (MPG-KPC 2) in left lateral (C) and anterior (D) view. E, F, half sacral centrum (MPG-KPC 16) in anterior (E) and ventral (F) view. Abbreviation: tp, broken basis of the transverse process. Scale bars are 5 cm (A-D) and 2 cm (E, F).
FIGURE 7. Anterior to mid-caudal vertebrae of Camarillasaurus cirugedae. A, B, partial anterior vertebral centrum MPG-KPC 21 in left lateral (A) and anterior (B) views. C, partial anterior caudal neural arch MPG-KPC 39 in left lateral view. D, E, partial anterior caudal centrum MPG-KPC 20 in ventral (D) and left lateral (E) view. F, G, partial mid-caudal vertebra MPZ 2022/182a in anterior (F) and left lateral (G; including the neural spine that belongs to this vertebra) view. Abbreviations: cf, chevron facets; poz, postzygapophysis; spdr, spinodiapophyseal ridge; tp, transverse process; vg, ventral groove. Scale bars are 5 cm.
FIGURE 8. Distal caudal vertebrae of Camarillasaurus cirugedae. A-E, vertebra MPG-KPC 11 in left lateral (A), anterior (B), posterior (C), dorsal (D), and ventral (E) view. F-J, vertebra MPG-KPC 10 in left lateral (F), anterior (G), posterior (H), dorsal (I), and ventral (J) view. K-O, vertebra MPG-KPC 12 in left lateral (K), anterior (L), posterior (M), dorsal (N), and ventral (O) view. P-T, vertebra MPG-KPC 13 in left lateral (P), anterior (Q), posterior (R), dorsal (S), and ventral (T) view. Abbreviations: bw, bony web; cf, chevron facets; ns, neural spine; poz, postzygapophysis; psr, prespinal ridge, tp, transverse process. Scale bars are 2 cm.
FIGURE 9. Ribs and chevrons of Camarillasaurus cirugedae. A, B, posterior dorsal rib of the left side, MPG-KPC 7, in posterior (A) and anterior (B) view. C, D, anterior chevron, MPG-KPC 5, in anterior (C) and right lateral (D) view. Abbreviations: ap, anterior process; ca, capitulum; fu, furrow; hc, haemal canal; su, sulcus; tu, tuberculum. Scale bars are 5 cm (A, B) and 2 cm (C, D).
FIGURE 10. Pectoral girdle elements of Camarillasaurus cirugedae. A-C, mid-shaft section of left scapula, MPG-KPC 30, in lateral (A), ventral (B), and dorsal (C) view. D, distal end of right scapula, MPG-KPC 1, in lateral view. Scale bars are 5 cm.
FIGURE 11. Possible anterior end of the preacetabular blade of the left ilium, MPG-KPC 23, in medial (A) and lateral (B) view. Scale bar is 5 cm.
FIGURE 12. Right femur of Camarillasaurus cirugedae, MPZ 2022/182c, in lateral (A), anterior (B), medial (C), posterior (D), distal (E), and proximal (F) view. Abbreviations: ag, articular groove; at, accessory trochanter; ctf, crista tibiofibularis; epc, epicondylar crest; ft, fourth trochanter; g, groove; lm, lateral mound; lt, lesser trochanter; mfe, m. femorotibialis externus. Scale bars are 5 cm.
FIGURE 13. Proximal end of right tibia of Camarillasaurus cirugedae, MPG-KPC 8, in lateral (A), medial (B), anterior (C), posterior (D), and proximal (E) view. Abbreviations: cc, cnemial crest; fc, fibular condyle; fcr, fibular crest. Scale bar is 5 cm.
FIGURE 14. Left pedal ungual four of Camarillasaurus cirugedae, MPZ 2022/182d, in proximal (A), right lateral (B), dorsal (C) and ventral (D) view. Abbreviations: cg, claw groove; d, depression; ft, flexor tubercle. Scale bar is 5 cm.
FIGURE 15. Phylogenetic hypothesis of tetanuran interrelationships, strict consensus tree resulting from a phylogenetic analysis using equal weights. See text for details of phylogenetic analysis.
FIGURE 16. Phylogenetic hypotheses of megalosauroid interrelationships. A, reduced consensus tree resulting from a phylogenetic analysis using equal weights. The different positions of taxa excluded by reduced consensus methods are marked by lowercase letters on the branches: a, Angaturama; b, Irritator; c, Piveteausaurus; e, Riojavenatrix; g, Sigilmassasaurus; h, MSNM V4047; i, Streptospondylus. B, reduced consensus tree resulting from a phylogenetic analysis using implied weights (k=12). The different positions of taxa excluded by reduced consensus methods are marked by lowercase letters on the branches: a, Angaturama; b, Duriavenator; c, Irritator; d, Magnosaurus; g, Riojavenatrix; h, MSNM V4047. See text for details of phylogenetic analysis.
