TABLE 1. Distribution of the Miocene-Pliocene elasmobranchs reported from Venice, Florida within the present-day Gulf of Mexico and Western North Atlantic Ocean. † = globally extinct species.
| Comparison of the Venice Elasmobranch Assemblage to Modern Assemblages in the USA | ||||
|---|---|---|---|---|
| Genus | Species | Venice Shelf, FL Peace River and Tamiami Fms., Miocene-Pliocene (This Study) | Present-day Gulf of Mexico (Castro, 2011; Last et al., 2016a; Carrier, 2017; Ebert et al., 2021) | Present-day Western North Atlantic Ocean (Bigelow and Schroeder, 1948; 1953; Castro, 2011; Last et al., 2016a; Ebert et al., 2021) |
| Squaliformes | ||||
| Squalus | sp. | X | X | X |
| † Isistius | triangulus | X | ||
| Heterodontiformes | ||||
| Heterodontus | sp. | X | Likely | |
| Orectolobiformes | ||||
| Ginglymostoma | cirratum | X | X | X |
| Lamniformes | ||||
| Carcharias | taurus | X | X | X |
| † Otodus | megalodon | X | ||
| † Parotodus | benedinii | X | ||
| Isurus | oxyrinchus | X | X | X |
| Carcharodon | carcharias | X | X | X |
| † Carcharodon | hastalis | X | ||
| Carcharhiniformes | ||||
| Scyliorhinus | sp. | X | X | X |
| Mustelus | sp. | X | X | X |
| Galeorhinus | sp. | X | ||
| † Hemipristis | serra | X | ||
| † Galeocerdo | aduncus | X | ||
| † Galeocerdo | mayumbensis | X | ||
| Galeocerdo | cuvier | X | X | X |
| † Physogaleus | contortus | X | ||
| Rhizoprionodon | sp. | X | X | X |
| Negaprion | brevirostris | X | X | X |
| Carcharhinus | cf. C. falciformis | X | X | X |
| Carcharhinus | leucas | X | X | X |
| Carcharhinus | obscurus | X | X | X |
| Carcharhinus | perezii | X | X | X |
| Carcharhinus | cf. C. brachyurus | X | ||
| Carcharhinus | cf. C. plumbeus | X | X | X |
| Carcharhinus | cf. C. altimus | X | X | X |
| Carcharhinus | limbatus | X | X | X |
| Carcharhinus | brevipinna | X | X | X |
| Carcharhinus | porosus | X | X | |
| Sphyrna | cf. S. zygaena | X | X | X |
| Sphyrna | cf. S. tiburo | X | X | X |
| Rhinopristiformes | ||||
| Rhynchobatus | sp. | X | ||
| Rhinobatos | sp. | X | ||
| Anoxypristis | sp. | X | ||
| Pristis | cf. P. pristis | X | X | |
| Rajiformes | ||||
| "Raja" | sp. | X | X | X |
| Myliobatiformes | ||||
| Hypanus | cf. H. americana | X | X | X |
| Hypanus | cf. H. rugosa | X | X | X |
| Rhinoptera | cf. R. bonasus | X | X | X |
| Mobula | cf. M. hypostoma | X | X | X |
| Mobula | cf. M. birostris | X | X | X |
| Aetomylaeus | sp. | X | ? | ? |
| Myliobatis | sp. | X | X | X |
| Aetobatus | cf. A. narinari | X | X | X |
TABLE 2. A comparison of the Venice Elasmobranch Assemblage with middle–late Miocene, Miocene–Pliocene, and Pliocene elasmobranch assemblages reported from Florida, USA. † = globally extinct species. A= Venice Shelf, Peace River and Tamiami Fms., Miocene–Pliocene (This Study), B= Torreya Fm., Middle Miocene, Gadsen County (Soto and MacFadden, 2014; Soto, 2016), C= Gainesville Creeks, Coosawhatchie Fm., Gainesville, Alachua County, Middle Miocene–Pliocene (Boyd, 2016), D= Love Bone Bed, Alachua Fm., Archer, Alachua County, Late Miocene (Webb et al., 1981), E= Montbrook Fossil Site (likely reworked Hawthorn Group), Williston, Levy County, Miocene–Pliocene (Ziegler et al., 2019), F= Manatee County Dam, Bone Valley Member, Peace River Fm., Bradenton, Manatee County, Pliocene (Webb and Tessman, 1968), G= Hickey Creek, Tamiami Fm., Alva, Lee County, Pliocene (Morgan and Pratt, 1983), H= Palmetto Fauna, Peace River Formation, Central Phosphate District, Polk, Hardee, Hillsborough, Manatee Counties, Miocene–Pliocene (Simons et al., 2014), I= Cookiecutter Creek, Tamiami Fm., North Port, Sarasota County, Pliocene (Perez and Marks, 2017). X*=Taxa identified to the genus level or as a similar species.
| Miocene-Pliocene Elasmobranch Assemblages in Florida, USA | ||||||||||
| Genus | Species | A | B | C | D | E | F | G | H | I |
| Squaliformes | ||||||||||
| Squalus | sp. | X | ||||||||
| † Isistius | triangulus | X | X | |||||||
| Heterodontiformes | ||||||||||
| Heterodontus | sp. | X | X | |||||||
| Orectolobiformes | ||||||||||
| Ginglymostoma | cirratum | X | X* | X* | X* | X* | X | |||
| Lamniformes | ||||||||||
| Carcharias | taurus | X | X* | X | X | X | X | |||
| † Otodus | megalodon | X | X | X | X | X | X | X | X | |
| † Parotodus | benedinii | X | ||||||||
| Isurus | oxyrinchus | X | X | X | X | |||||
| Carcharodon | carcharias | X | X | |||||||
| † Carcharodon | hastalis | X | X | X | X | X | X | X | X | X |
| Carcharhiniformes | ||||||||||
| Scyliorhinus | sp. | X | ||||||||
| Mustelus | sp. | X | ||||||||
| Galeorhinus | sp. | X | X | |||||||
| † Hemipristis | serra | X | X | X | X | X | X | X | X | X |
| † Galeocerdo | aduncus | X | X | X | X | |||||
| † Galeocerdo | mayumbensis | X | X | X | ||||||
| Galeocerdo | cuvier | X | X | X | X | X | X | X | ||
| † Physogaleus | contortus | X | X | X | X | X | X | X | ||
| Rhizoprionodon | sp. | X | X | X | X | X | X | X | X | X |
| Negaprion | brevirostris | X | X | X | X | X | X | X | ||
| Carcharhinus | cf. C. falciformis | X | X | X | X | |||||
| Carcharhinus | leucas | X | X* | X | X | X | X | X | X | |
| Carcharhinus | obscurus | X | ||||||||
| Carcharhinus | perezii | X | X | X | ||||||
| Carcharhinus | cf. C. brachyurus | X | X | X | ||||||
| Carcharhinus | cf. C. plumbeus | X | X | X | ||||||
| Carcharhinus | cf. C. altimus | X | ||||||||
| Carcharhinus | limbatus | X | X | X | ||||||
| Carcharhinus | brevipinna | X | X | |||||||
| Carcharhinus | porosus | X | ||||||||
| Sphyrna | cf. S. zygaena | X | X | X* | X* | X* | ||||
| Sphyrna | cf. S. tiburo | X | ||||||||
| Rhinopristiformes | ||||||||||
| Rhynchobatus | sp. | X | X | X | X | |||||
| Rhinobatos | sp. | X | X | X | X | |||||
| Anoxypristis | sp. | X | X | X | X | |||||
| Pristis | cf. P. pristis | X | X* | X | X | |||||
| Rajiformes | ||||||||||
| "Raja" | sp. | X | X | |||||||
| Myliobatiformes | ||||||||||
| Hypanus | cf. H. americana | X | X | X | X | X | X | |||
| Hypanus | cf. H. say | X | ||||||||
| Rhinoptera | cf. R. bonasus | X | X | X | X | X | ||||
| Mobula | cf. M. hypostoma | X | X | |||||||
| Mobula | cf. M. birostris | X | ||||||||
| Aetomylaeus | sp. | X | X | X | ||||||
| Myliobatis | sp. | X | X | X | X | X | X | X | ||
| Aetobatus | cf. A. narinari | X | X | X | ||||||
TABLE 3. A comparison of the Venice Elasmobranch Assemblage with other Miocene and Early Pliocene elasmobranch assemblages reported from the USA. For Onslow Bay and the Nutrien Mine: M = Miocene; P = Pliocene; * = identified to only the genus level or different species name utilized; † = globally extinct species.
| Miocene-Pliocene Elasmobranch Assemblages ACP, USA | ||||||
| Genus | Species | Venice Shelf, FL Peace River and Tamiami Fms., Miocene-Pliocene (This Study) | Onslow Bay, NC Pungo River and Yorktown Fms. Early Miocene-Pliocene (Maisch et al., 2018) | Nutrien Mine, NC Pungo River and Yorktown Fms. Early Miocene-Pliocene (Purdy et al., 2001) | Carmel Church Quarry, VA Calvert Fm. Middle Miocene (Hastings and Dooley, 2017) | Calvert Cliffs, MD Calvert, Choptank, and St. Marys Fms. Late Early-Late Miocene (Kent, 2018) |
| Squaliformes | ||||||
| Squalus | sp. | X | XP | X | ||
| † Isistius | triangulus | X | X * | |||
| Heterodontiformes | ||||||
| Heterodontus | sp. | X | ||||
| Orectolobiformes | ||||||
| Ginglymostoma | cirratum | X | XM* | |||
| Lamniformes | ||||||
| Carcharias | taurus | X | XP | XP | X | X |
| † Otodus | megalodon | X | XP | XMP | X | X |
| † Parotodus | benedinii | X | XP | XP | X | |
| Isurus | oxyrinchus | X | XMP | XMP | X | |
| Carcharodon | carcharias | X | XP | XP | X | |
| † Carcharodon | hastalis | X | XP | XP | X | X |
| Carcharhiniformes | ||||||
| Scyliorhinus | sp. | X | XMP | X | ||
| Mustelus | sp. | X | XM | X | ||
| Galeorhinus | sp. | X | XMP | |||
| † Hemipristis | serra | X | XMP | XMP | X | X |
| † Galeocerdo | aduncus | X | XM | XM* | X * | X |
| † Galeocerdo | mayumbensis | X | ||||
| Galeocerdo | cuvier | X | XP | XP | ||
| † Physogaleus | contortus | X | XM | XM | X | X |
| Rhizoprionodon | sp. | X | XMP | XM | X | |
| Negaprion | brevirostris | X | XP | XMP | X * | X* |
| Carcharhinus | cf. C. falciformis | X | XMP | XM | X | |
| Carcharhinus | leucas | X | XMP | X * | X | |
| Carcharhinus | obscurus | X | XP | |||
| Carcharhinus | perezii | X | XMP | X | ||
| Carcharhinus | cf. C. brachyurus | X | XM | |||
| Carcharhinus | cf. C. plumbeus | X | XMP | X | ||
| Carcharhinus | cf. C. altimus | X | X | |||
| Carcharhinus | limbatus | X | ||||
| Carcharhinus | brevipinna | X | ||||
| Carcharhinus | porosus | X | ||||
| Sphyrna | cf. S. zygaena | X | XMP | X* | X* | |
| Sphyrna | cf. S. tiburo | X | ||||
| Rhinopristiformes | ||||||
| Rhynchobatus | sp. | X | X | |||
| Rhinobatos | sp. | X | XM | |||
| Anoxypristis | sp. | X | XP | X* | ||
| Pristis | cf. P. pristis | X | XP * | |||
| Rajiformes | ||||||
| "Raja" | sp. | X | XP | X | ||
| Myliobatiformes | ||||||
| Hypanus | cf. H. americana | X | XM | |||
| Hypanus | cf. H. say | X | XMP* | X | ||
| Rhinoptera | cf. R. bonasus | X | XM* | X* | ||
| Mobula | cf. M. hypostoma | X | MP* | X* | ||
| Mobula | cf. M. birostris | X | ||||
| Aetomylaeus | sp. | X | X * | X * | ||
| Myliobatis | sp. | X | X | |||
| Aetobatus | cf. A. narinari | X | XMP * | X | X * | |
TABLE 4. A comparison of the Venice Elasmobranch Assemblage with late Miocene, Miocene–Pliocene, and Pliocene elasmobranch assemblages from Central and South America. Note that taxonomic diversity is highly variable among assemblages, some species names have since been revised, and diversity differences may stem from taxonomic splitting and lumping. † = globally extinct species. A= Peace River and Tamiami Fms., Miocene– Pliocene, FL (This Study), B= Chagres Fm., Late Miocene, Panama (Carrillo-Briceño et al., 2015), C= Gatun and Chucunaque Fms., Late Miocene, Panama (Pimiento et al., 2013; Perez et al., 2017), D= Curré Fm., Late Miocene, Costa Rica (Laurito and Valerio, 2008), E= Pisco Fm., Late Miocene, Peru (Landini et al., 2017a,b), F= Uscari Fm., Miocene– Pliocene, Costa Rica (Laurito, 1999; Laurito and Valerio, 2021), G= Onzole–Bahia Fms., Late Miocene– Pliocene, Ecuador (Longbottom, 1979; Carrillo-Briceño et al., 2014), H= Tirabuźon Fm., Pliocene, Mexico (Lira-Beltrán et al., 2020), I= Urumaco Sequence (Soccorro, Urumaco, and Codore Fms.), Miocene–Pliocene, Venezuela (Carrillo-Briceño et al., 2015), J= Horcón Fm., Pliocene, Chile (Carrillo-Briceño et al., 2013). X*=Taxa identified to the genus level or as a similar species.
| Miocene-Pliocene Elasmobranch Assemblages in Central and South America | |||||||||||
| Genus | Species | A | B | C | D | E | F | G | H | I | J |
| Squaliformes | |||||||||||
| Squalus | sp. | X | X | X | X | ||||||
| † Isistius | triangulus | X | X* | X | X | ||||||
| Heterodontiformes | |||||||||||
| Heterodontus | sp. | X | X* | X* | X* | X* | |||||
| Orectolobiformes | |||||||||||
| Ginglymostoma | cirratum | X | X* | ||||||||
| Lamniformes | |||||||||||
| Carcharias | taurus | X | X* | X | X* | ||||||
| † Otodus | megalodon | X | X | X | X | X | X | X | X | ||
| † Parotodus | benedinii | X | X | ||||||||
| Isurus | oxyrinchus | X | X | X* | X | X* | X | ||||
| Carcharodon | carcharias | X | X | X | |||||||
| † Carcharodon | hastalis | X | X | X | |||||||
| Carcharhiniformes | |||||||||||
| Scyliorhinus | sp. | X | X | ||||||||
| Mustelus | sp. | X | X | X | X* | X | |||||
| Galeorhinus | sp. | X | X* | X* | X* | ||||||
| † Hemipristis | serra | X | X | X | X | X | X | X | X | ||
| † Galeocerdo | aduncus | X | X | X | X | ||||||
| † Galeocerdo | mayumbensis | X | |||||||||
| Galeocerdo | cuvier | X | X | X | X | X | |||||
| † Physogaleus | contortus | X | X | X | |||||||
| Rhizoprionodon | sp. | X | X | X | X* | X | X* | X | |||
| Negaprion | brevirostris | X | X | X | X | X | X | ||||
| Carcharhinus | cf. C. falciformis | X | X | X | |||||||
| Carcharhinus | leucas | X | X | X | X* | X | X* | X* | X | X | |
| Carcharhinus | obscurus | X | X | X | X | X | |||||
| Carcharhinus | perezii | X | X | ||||||||
| Carcharhinus | cf. C. brachyurus | X | X | X | X | X | |||||
| Carcharhinus | cf. C. plumbeus | X | X | X | X | ||||||
| Carcharhinus | cf. C. altimus | X | |||||||||
| Carcharhinus | limbatus | X | X | ||||||||
| Carcharhinus | brevipinna | X | X | X | |||||||
| Carcharhinus | porosus | X | X | ||||||||
| Sphyrna | cf. S. zygaena | X | X* | X | X* | X | X | X* | X | X | |
| Sphyrna | cf. S. tiburo | X | |||||||||
| Rhinopristiformes | |||||||||||
| Rhynchobatus | sp. | X | X | X* | X | ||||||
| Rhinobatos | sp. | X | X | X | |||||||
| Anoxypristis | sp. | X | X* | ||||||||
| Pristis | cf. P. pristis | X | X | X* | |||||||
| Rajiformes | |||||||||||
| "Raja" | sp. | X | X* | X | X* | ||||||
| Myliobatiformes | |||||||||||
| Hypanus | cf. H. americana | X | X* | X* | X* | X* | X* | X* | X* | ||
| Hypanus | cf. H. say | X | |||||||||
| Rhinoptera | cf. R. bonasus | X | X* | X* | X* | X* | X* | ||||
| Mobula | cf. M. hypostoma | X | X* | X | X* | X* | |||||
| Mobula | cf. M. birostris | X | X | ||||||||
| Aetomylaeus | sp. | X | X* | ||||||||
| Myliobatis | sp. | X | X | X* | X | X | X | X | X | X* | |
| Aetobatus | cf. A. narinari | X | X* | X* | X* | X* | X* | ||||
TABLE 5. Global Miocene-Pliocene occurrence of the Venice elasmobranch taxa. † = globally extinct species. References utilized: Atlantic Coastal Plain and Gulf Coastal Plain, USA (Kimmel and Purdy, 1984; Purdy et al., 2001; Boessenecker et al., 2018; Kent, 2018; Maisch et al., 2018; Perez, 2022); Western North America (Boessenecker, 2011, 2016; Boessenecker et al., 2014; Lira-Beltrán et al., 2020); Caribbean, Central, and South America (Davies, 1964; Gillette, 1984; Applegate, 1986; Long, 1993; Iturralde-Vinent et al., 1996; Flemming and McFarlane, 1998; Nieves-Rivera et al., 2003; Laurito Mora, 1999, 2004; Apolín et al., 2004; Portell et al., 2008; Cione et al., 2011, 2012; Ehret et al., 2012; Pimiento et al., 2013; Collareta et al., 2017a,2017b, 2017c; Landini et al., 2017a; MacFadden et al., 2017; Perez et al., 2017; Carrillo-Briceño et al., 2014, 2015a, 2015b, 2016a, 2016b, 2018, 2019; Laurito Mora and Valerio, 2008, 2021; Laurito Mora et al., 2022); Africa (Cook et al., 2010; Ávila et al., 2012; Pawellek et al., 2012; Govender and Chinsamy, 2013; Andrianavalona et al., 2015; Betancort et al., 2016); Europe (Holec et al., 1995; Antunes et al., 1999; Antunes and Balbino, 2004; Cappetta and Cavallo, 2006; Carnevale et al., 2006; Kocsis, 2007; Marsili, 2007; Marsili et al., 2007; Cigala-Fulgosi et al., 2009; Vialle et al., 2011; Cappetta, 2012; Reinecke and Radwánski, 2015; Szabó and Kocsis, 2016; Fialho et al., 2019, 2021; Collareta et al., 2021; Szabó et al., 2021, 2023); Asia (Itoigawa and Nishimoto, 1974; Uyeno, 1978; Sahni and Mehrotra, 1981; Goto et al., 1984; Shimada, 1987; Sharma and Patniak, 2013, 2014; Kocsis et al., 2018; Razak and Kocsis, 2018; Sharma et al., 2021); Australia (Pledge, 1967; Kemp, 1991).
| Subcontinent/Continent | ACP, GCP | WNA | Carib., CA, SA | Africa | Europe | Asia | Australia | |
|---|---|---|---|---|---|---|---|---|
| Squaliformes | ||||||||
| Squalus | sp. | X | X | X | X | X | ||
| † Isistius | triangulus | X | X | X | ||||
| Heterodontiformes | ||||||||
| Heterodontus | spp. | X | X | X | X | X | ||
| Orectolobiformes | ||||||||
| Ginglymostoma | cirratum | X | X | X | ||||
| Lamniformes | ||||||||
| Carcharias | taurus | X | X | X | X | X | X | |
| † Otodus | megalodon | X | X | X | X | X | X | X |
| † Parotodus | benedinii | X | X | X | X | X | X | |
| Isurus | oxyrinchus | X | X | X | X | X | X | X |
| Carcharodon | carcharias | X | X | X | X | X | X | X |
| † Carcharodon | hastalis | X | X | X | X | X | X | X |
| Carcharhiniformes | ||||||||
| Scyliorhinus | sp. | X | X | X | X | |||
| Mustelus | sp. | X | X | X | X | X | ||
| Galeorhinus | sp. | X | X | X | X | |||
| † Hemipristis | serra | X | X | X | X | X | X | |
| † Galeocerdo | aduncus | X | X | X | X | X | X | |
| † Galeocerdo | mayumbensis | X | X | X | X | |||
| Galeocerdo | cuvier | X | X | X | X | X | X | X |
| † Physogaleus | contortus | X | X | X | X | X | ||
| Rhizoprionodon | spp. | X | X | X | X | X | X | X |
| Negaprion | brevirostris | X | X | X | X | sp. | X | |
| Carcharhinus | cf. C. falciformis | X | X | X | X | X | ||
| Carcharhinus | leucas | X | X | X | X | X | X | |
| Carcharhinus | obscurus | X | X | X | X | X | ||
| Carcharhinus | perezii | X | X | X | ||||
| Carcharhinus | cf. C. brachyurus | X | X | X | X | X | X | |
| Carcharhinus | cf. C. plumbeus | X | X | X | X | |||
| Carcharhinus | cf. C. altimus | X | ||||||
| Carcharhinus | cf. C. limbatus | X | X | X | ||||
| Carcharhinus | cf. C. brevipinna | X | X | X | ||||
| Carcharhinus | cf. C. porosus | X | X | |||||
| Sphyrna | spp. | X | X | X | X | X | X | X |
| Sphyrna | cf. S. zygaena | X | X | X | X | X | ||
| Sphyrna | cf. S. tiburo | X | ||||||
| Rhinopristiformes | ||||||||
| Rhynchobatus | sp. | X | X | X | X | X | ||
| Rhinobatos | sp. | X | X | X | X | X | X | |
| Anoxypristis | sp. | X | X | X | X | X | X | |
| Pristis | cf. P. pristis | X | X | X | X | |||
| Rajiformes | ||||||||
| "Raja" | spp. | X | X | X | X | X | ||
| Myliobatiformes | ||||||||
| Hypanus | spp. | X | X | X | X | X | X | |
| Rhinoptera | cf. R. bonasus | X | X | X | X | X | ||
| Mobula | cf. M. hypostoma | X | X | X | ||||
| Mobula | cf. M. birostris | X | X | |||||
| Aetomylaeus | spp. | X | X (as Aetobatus) |
X | X | X | ||
| Myliobatis | spp. | X | X | X | X | X | X | X |
| Aetobatus | cf. A. narinari | X | X | X | X | X | ||
