TABLE 1. Measurements taken on distal humeri in this study. Columns (from left to right) are: measurements (abbreviation and name); measurement description; taxon of study; and source reference for each measurement. We took the following measurements (Figure 2) on all multituberculate and therian specimens, contingent upon specimen completeness (see Table 2). The anatomical view from which the measurement was taken is indicated in parentheses in the description, as it relates to either multituberculate (Mu) or therian (Th) humeri, see text for more details. Measurements are identical to those in the indicated sources, with the following exceptions: our measurement TDW is ‘BW’ of Szalay and Dagosto (1980) and measurement 5 of Argot (2001); NCW is measurement 6 of Argot (2001) and ‘EEC’ of Boyer et al. (2010); TLW is ‘TLE’ of Szalay and Dagosto (1980); measurements TD and CD are modified from Szalay and Dagosto (1980) and Boyer et al. (2010). Multituberculate measurements lacking references are based on the structurally (but not necessarily functional) analogous measurement for therian humeri (see Figure 2).
Measurements | Description | Taxon | Source | |
TDW | Total Distal Width | Medial edge of specimen to lateral edge, measured at the maximum extensions of the entepicondyle and ectepicondyle; parallel to other distal width measures (anterior view in therians; ventral view in multituberculates) | Th, Mu | Deischl 1964; Szalay and Dagosto 1980; Argot 2001 |
AW | Articular Surface Width | Eu: Lateral edge of capitulum to medial edge of trochlea (anterior)
Mu: Lateral edge of radial condyle to medial edge of ulnar condyle (ventral) |
Th, Mu | Szalay and Dagosto 1980; Szalay and Sargis 2001 |
TLW | Total Lateral Width | Eu: Medial edge of trochlea to lateral edge of ectepicondyle (anterior)
Mu: Medial edge of ulnar condyle to lateral edge of ectepicondyle (ventral) |
Th, Mu | Szalay and Dagosto 1980 |
NCW | Entepicondylar Width | Eu: Medial edge of trochlear lip to medial edge of entepicondyle (anterior)
Mu: From midpoint of groove between entepicondyle and ulnar condyle to medial edge of entepicondyle (ventral) |
Th, Mu | Argot 2001; Boyer et al. 2010 |
TW | Trochlear Width | Medial edge of trochlea (trochlear lip) to lateral edge of trochlea (medial edge of intercondylar groove) (anterior) | Th | Szalay and Dagosto 1980; Boyer et al. 2010 |
CW | Capitulum Width | Lateral edge of capitulum to medial edge of intercondylar groove (anterior) | Th | Szalay and Dagosto 1980; Boyer et al. 2010 |
TL | Trochlear Length | Length from proximal margin to distal margin of trochlear lip (anterior) | Th | Szalay and Dagosto 1980; Boyer et al. 2010 |
CL | Capitulum Length | Length from proximal margin to distal margin of capitulum (anterior) | Th | this study |
TD | Trochlear Depth | Maximum length from anterior surface to posterior surface of trochlea, parallel to CD (distal) | Th | Szalay and Dagosto 1980; Boyer et al. 2010 |
CD | Capitulum Depth | Length of capitulum articular surface, from anterior to posterior extent, parallel to TD (distal) | Th | Szalay and Dagosto 1980; Boyer et al. 2010 |
UCW | Ulnar Condyle Width | Medial edge of ulnar condyle to lateral edge of ulnar condyle (ventral) | Mu | this study |
RCW | Radial Condyle Width | Medial edge of radial condyle to lateral edge of radial condyle (ventral) | Mu | this study |
UCL | Ulnar Condyle Length | Length from proximal margin to distal margin of ulnar condyle (ventral) | Mu | this study |
RCL | Radial Condyle Length | Length from proximal margin to distal margin of radial condyle (ventral) | Mu | this study |
UCD | Ulnar Condyle Depth | Maximum length from ventral surface to dorsal surface ulnar condyle, parallel to RCD (distal) | Mu | this study |
RCD | Radial Condyle Depth | Maximum length from ventral surface to dorsal surface radial condyle, parallel to UCD (distal) | Mu | this study |
TABLE 2. Linear measurements of multituberculate distal humeri specimens. See Table 1 for measurement descriptions; all measurements on multituberculate humeri are taken in ventral view except for RCD and UCD, which are taken in distal view. All measurements are in millimeters. Abbreviations: Fm, Formation; HC, Hell Creek Formation; La, Lancian Biozone; Lan, Lance Formation; Morph., Morphotype; Pu1, Puercan 1 Biozone; Pu3, Puercan 3 Biozone; Tu, Tullock Formation. Asterisks indicate minimum size values, where preserved morphology was measured but the full measurement was not possible, due to specimen breakage; “-” indicates specimen was too broken for measurement; “?” indicates more tentative assignment to morphotype, due to specimen breakage. Locality numbers follow the system of each institution; those beginning with “V” are UCMP localities; those beginning with “C” are UWBM localities (see Appendix 1 for additional locality details).
Specimen | Locality | Fm | Biozone | Morph. | TDW | AW | TDW | NCW | UCW | RCW | UCL | RCL | UCD | RCD |
195999 | V5620 | Lan | La | MuA | 3.695 | 2.060 | 2.550 | 1.045 | 0.950 | 1.115 | 1.365 | 1.195 | 1.625 | 1.525 |
195974 | V65127 | Tu | BCA | MuA | 3.995 | 2.005 | 2.455 | 1.255 | 0.765 | 1.000 | 1.305 | 1.140 | 1.530 | 1.510 |
195982 | V65127 | Tu | BCA | MuA | - | 2.040 | 2.210 | - | 0.865 | 1.170 | 1.405 | 1.055 | 1.445 | - |
122045 | V70201 | Tu | BCA | MuA | 4.120 | 2.105 | 2.565 | 1.375 | 0.875 | 1.235 | 1.315 | 1.165 | 1.650 | 1.580 |
196956 | V70201 | Tu | BCA | MuA | - | 1.960* | - | - | 0.780 | 1.120* | 1.225 | 1.005 | 1.275 | - |
196957 | V70201 | Tu | BCA | MuA | 3.915 | 2.090 | 2.615 | 1.175 | 0.840 | 1.160 | 1.465 | 1.240 | 1.460 | 1.455 |
70976 | C0338 | Tu | BCA | MuA | - | 1.850 | 2.420 | - | 0.855 | 1.070 | 1.345 | 1.150 | 1.440 | 1.400 |
70977 | C0338 | Tu | BCA | MuA | 3.695 | - | 2.580 | 1.075 | 1.065 | - | 1.480 | - | 1.415 | - |
195998 | V5620 | Lan | La | ?MuA | - | - | - | - | 0.690 | - | 0.805 | - | 1.805 | - |
153016 | V74111 | Tu | Pu1 | ?MuA | - | 2.165* | - | - | - | 1.240 | 1.475* | 1.240* | 1.585 | 1.475* |
195933 | V99438 | Tu | Pu3 | MuB | - | - | - | 1.240 | 1.080 | - | 1.465 | - | 1.670 | - |
174490 | V73087 | HC | La | MuC | - | 3.555* | - | - | 1.690 | 2.055 | 2.440 | 1.980 | 2.845 | 2.445 |
195997 | V5620 | Lan | La | MuC | - | 2.525 | 3.150 | - | 0.985 | 1.360 | 1.925 | 1.600 | 2.145 | 2.000 |
174420 | V70209 | Tu | BCA | MuC | 5.290 | 2.975 | 3.595 | 1.480 | 1.295 | 1.630 | 1.975 | 2.030* | 2.130 | 1.975 |
153039 | V84193 | HC | Pu1 | MuC | 6.460 | 3.090 | 4.250 | 1.960 | 1.570 | 1.700 | 2.185 | 1.830 | 2.555 | 2.335 |
197687 | V74111 | Tu | Pu1 | MuC | - | 3.565* | - | - | 1.480* | 2.050* | 2.240 | 1.885 | 2.330 | - |
195932 | V99438 | Tu | Pu3 | MuC | 5.350* | 3.035 | 3.820 | 1.460* | 1.505 | 1.640 | 2.180 | 1.765 | 2.515 | 2.295 |
195945 | V74122 | Tu | Pu3 | MuC | 5.590* | 3.040 | 3.815 | 1.480 | 1.270 | 1.420 | 2.455 | 1.755 | 2.345 | 2.305 |
195990 | V5620 | Lan | La | MuD | 7.310* | 4.230* | 5.470* | 1.835* | - | 2.160 | - | 2.135 | - | 2.350 |
174439 | V70209 | Tu | BCA | MuE | - | - | - | 2.385 | 1.855 | - | 3.265 | - | 3.200 | - |
174442 | V70209 | Tu | BCA | MuE | - | - | - | - | 1.965 | - | 2.730* | - | 2.600* | 2.680 |
195973 | V65127 | Tu | BCA | MuE | 8.130 | 4.025 | 5.080 | 2.445 | 1.485 | 2.410 | 2.810 | 2.265 | 3.200 | 3.145 |
97031 | C1845 | Tu | BCA | MuE | 8.420 | 4.400 | 5.230 | 2.595 | 2.000 | 2.565 | 2.695 | 2.485 | 3.355 | 3.335 |
174412 | V84162 | HC | Pu1 | MuE | - | - | - | - | 2.360 | - | 3.535 | - | 3.910 | - |
127384 | V87001 | HC | La | MuF | 14.025 | 7.730 | 9.990 | 3.970 | 3.865 | 4.435 | 5.440 | 5.095 | 6.420 | 5.580 |
174400 | V85092 | HC | La | MuF | - | 6.900 | - | - | 3.135 | 3.945 | 4.985 | 4.260 | 5.180 | 4.530 |
174232 | V72207 | HC | La | MuF | - | - | - | - | - | 5.050 | - | 5.655 | - | 6.365 |
174225 | V72207 | HC | La | MuF | - | 6.815* | - | 3.255 | 3.435 | 4.215* | 5.045 | 4.900 | 5.835 | - |
174218 | V73097 | HC | La | MuF | - | 7.820* | - | - | 2.450 | 4.325 | 5.605 | 5.145 | 6.420 | - |
101970 | C1521 | HC | La | MuF | 13.705 | 6.945 | 9.105 | 3.995 | 2.985 | 3.915 | 5.030 | 3.900 | 5.030 | 4.405 |
TABLE 3. Linear measurements of therian distal humeri specimens. See Table 1 for measurement descriptions; all measurements on therian humeri are taken in anterior view except for CD and TD, which are taken in distal view . All measurements are in millimeters. Abbreviations: Fm, Formation; HC, Hell Creek Formation; La, Lancian Biozone; Morph., Morphotype; Pu1, Puercan 1 Biozone; Pu3, Puercan 3 Biozone; Tu, Tullock Formation. Asterisks indicate minimum size values, where preserved morphology was measured but the full measurement was not possible, due to specimen breakage; “-” indicates specimen was broken; “?” indicates more tentative assignment to morphotype; “‡” indicates specimen was used in 2D GM locomotor analyses.Locality numbers follow the system of each institution: those beginning with “V” are UCMP localities; those beginning with “C” are UWBM localities (see Appendix 1 for additional locality details). Five additional specimens from the BCA biozone (UCMP 151945, 151965, 151983, and 151984 from loc. V65127, and UCMP 174429 from loc. V70209) are tentatively attributed to EuD but are not listed below as they lack preservation sufficient for measurement, see text for more details.
Specimen | Locality | Fm | Biozone | Morph. | TDW | AW | TLW | NCW | TW | CW | TL | CL | TD | CD |
127391 | V88007 | HC | La | ThA | 4.900* | - | - | 2.380 | 1.210 | - | 1.340 | - | 1.380 | - |
152394 | V93169 | HC | La | MeA | 18.235* | - | - | - | - | - | - | - | - | - |
153100 | V99438 | Tu | Pu3 | EuA | 6.400* | 3.850 | 3.850 | 2.900* | 1.330 | 2.406 | 1.630 | 1.579 | 1.756 | 1.920 |
97114‡ | C1115 | HC | La | EuB | 8.53* | 4.765 | 5.120 | 3.400* | 1.705 | 2.895 | 1.960 | 1.945 | 2.355 | 2.335* |
151991‡ | V71203 | HC | BCA | EuC | 9.520 | 5.635 | 5.735 | 3.820 | 1.670 | 3.945 | 2.680 | 2.515 | 3.030 | 3.310 |
153023 | V74111 | Tu | Pu1 | EuC | - | - | - | 4.015 | 1.510* | - | 2.500* | - | 3.160 | - |
151985 | V70201 | Tu | BCA | EuD | 7.810* | 5.295 | 5.285 | - | 1.890 | 3.255 | 2.175 | .720 | 2.235 | 2.495 |
151944 | V65127 | Tu | BCA | EuD | 8.845* | 5.735 | 6.005 | 2.190* | 2.185 | 3.655 | 1.940 | 2.320 | 2.230* | 2.575* |
151946 | V65127 | Tu | BCA | EuD | 8.800* | 5.270 | 6.060 | - | 1.970 | 3.255 | 1.665 | 2.080 | 2.185 | 2.510* |
151964‡ | V65127 | Tu | BCA | EuD | 9.225 | 5.340 | 5.700 | 3.550 | 1.750 | 3.585 | 1.645 | 2.005 | 2.225 | 2.380 |
153036‡ | V65127 | Tu | BCA | EuD | 9.935 | 5.675 | 5.850 | 4.210 | 1.820 | 3.800 | 1.775 | 2.155 | 2.470 | 2.710 |
218901 | V91065 | Tu | Pu1 | EuE | 17.925* | 10.685 | 10.850 | 7.175* | 2.950 | 7.620 | 4.575 | 5.110 | 6.280 | 7.525 |
92928 | C1163 | Tu | Pu3 | EuE | 15.030* | 9.550 | 9.700 | - | 2.640 | 6.960 | 4.330 | 4.580 | 5.120* | 5.320 |
153089 | V72129 | Tu | Pu3 | EuF | 19.415* | 17.280 | 17.395 | - | 5.480 | 11.820 | 6.945 | 7.280 | 8.735* | 9.670 |
192678 | V72124 | Tu | Pu3 | EuG | - | - | - | 10.300 | 5.095 | - | 7.435 | - | 9.425* | - |
TABLE 4. Modern mammalian specimens used in geometric morphometrics analyses. Species list and locomotor references are modified from Chen and Wilson 2015; see text for more details. All specimens are from the UWBM except for those with specimen numbers A2001 and S2008, which are taken from images of specimens in distal view in Argot (2001, figure 8) and Salton and Sargis (2008, figure 4.10), respectively. Code number assigned (and table sorted) as follows: clade, order, family, and locomotor mode. Locomotor mode abbreviations are as follows: A, arboreal; F, fossorial; G, gliding; S, saltatorial; Sa, semiaquatic; Sc, scansorial; Sf, semifossorial; T, terrestrial. Other abbreviations: F, female; LM, locomotor mode; M, male; Spec., specimen number; U, sex unknown.
Clade | Order | Code | LM | Genus | Species | Family | Spec. | Sex | Locomotor Reference |
Metatheria | |||||||||
Dasyuromorphia | |||||||||
1 | T | Antechinus | swainsonii | Dasyuridae | 68900 | M | Nowak 1999 | ||
2 | T | Sarcophilus | harrisii | Dasyuridae | 20671 | M | Van Valkenburgh 1987 | ||
Didelphimorphia | |||||||||
3 | A | Caluromys | philander | Didelphidae | A2001 | U | Argot 2001; Nowak 1999 | ||
4 | A | Caluromys | derbianus | Didelphidae | 32255 | U | Argot 2001 | ||
5 | A | Micoureus | demerarae | Didelphidae | A2001 | U | Argot 2001 | ||
6 | Sc | Didelphis | virginiana | Didelphidae | 35525 | F | Argot 2001 | ||
7 | Sc | Didelphis | virginiana | Didelphidae | 74661 | M | Argot 2001 | ||
8 | Sc | Didelphis | marsupialis | Didelphidae | A2001 | U | Argot 2001; Nowak 1999 | ||
9 | Sc | Marmosa | murina | Didelphidae | A2001 | U | Argot 2001; Nowak 1999 | ||
10 | T | Metachirus | nudicaudatus | Didelphidae | 35438 | F | Argot 2001; Delciellos and Vieira 2006 | ||
11 | T | Monodelphis | brevicaudata | Didelphidae | A2001 | U | Argot 2001; Nowak 1999 | ||
Diprotodontia | |||||||||
12 | G | Petaurus | breviceps | Petauridae | 34181 | M | Smith 1973; Nowak 1999; Körtner and Geiser 2000 | ||
13 | G | Petaurus | breviceps | Petauridae | 72928 | U | Smith 1973; Nowak 1999; Körtner and Geiser 2000 | ||
14 | A | Trichosurus | vulpecula | Phalangeridae | 68913 | M | Nowak 1999 | ||
15 | S | Potorous | tridactylus | Potoroidae | 41025 | F | Bassarova et al. 2009 | ||
16 | S | Potorous | tridactylus | Potoroidae | 34200 | M | Bassarova et al. 2009 | ||
17 | S | Aepyprymnus | rufescens | Potoroidae | 68898 | F | Bassarova et al. 2009 | ||
18 | A | Pseudocheirus | peregrinus | Pseudocheiridae | 68924 | F | Bassarova et al. 2009 | ||
19 | A | Pseudocheirus | peregrinus | Pseudocheiridae | 68908 | M | Bassarova et al. 2009 | ||
Peramelemorphia | |||||||||
20 | T | Perameles | nasuta | Peramelidae | 68919 | F | Nowak 1999 | ||
Eutheria | |||||||||
Afrosoricida | |||||||||
21 | Sa | Potamogale | velox | Potamogalidae | S2008 | U | Nowak 1999; Salton and Sargis 2008 | ||
22 | A | Dendrohyrax | arboreus | Procaviidae | 39039 | F | Nowak 1999 | ||
23 | A | Dendrohyrax | arboreus | Procaviidae | 39038 | M | Nowak 1999 | ||
24 | Sa | Limnogale | mergulus | Tenrecidae | S2008 | U | Nowak 1999; Salton and Sargis 2008 | ||
25 | Sc | Echinops | telfairi | Tenrecidae | 34168 | F | Endo et al. 2006; Salton and Sargis 2008 | ||
26 | Sc | Echinops | telfairi | Tenrecidae | S2008 | U | Endo et al. 2006; Salton and Sargis 2008 | ||
27 | Sf | Hemicentetes | semispinosus | Tenrecidae | S2008 | U | Nowak 1999; Salton and Sargis 2008 | ||
28 | Sf | Oryzorictes | sp. | Tenrecidae | S2008 | U | Nowak 1999; Salton and Sargis 2008 | ||
29 | T | Setifer | setosus | Tenrecidae | S2008 | U | Salton and Sargis 2008 | ||
30 | T | Tenrec | ecaudatus | Tenrecidae | S2008 | U | Salton and Sargis 2008 | ||
31 | T | Microgale | cowani | Tenrecidae | S2008 | U | Nowak 1999; Salton and Sargis 2008 | ||
32 | T | Microgale | dobsoni | Tenrecidae | S2008 | U | Nowak 1999; Salton and Sargis 2008 | ||
33 | T | Microgale | talazaci | Tenrecidae | S2008 | U | Nowak 1999; Salton and Sargis 2008 | ||
Carnivora | |||||||||
34 | T | Vulpes | vulpes | Canidae | 39490 | F | Lariviére and Pasitschniak-Arts 1996; Meachen-Samuels 2010 | ||
35 | T | Vulpes | vulpes | Canidae | 39489 | M | Lariviére and Pasitschniak-Arts 1996; Meachen-Samuels 2010 | ||
36 | Sc | Urocyon | cinereoargenteus | Felidae | 77676 | F | Van Valkenburgh 1987; Meachen-Samuels 2010 | ||
37 | Sc | Urocyon | cinereoargenteus | Felidae | 35221 | M | Van Valkenburgh 1987; Meachen-Samuels 2010 | ||
38 | Sc | Suricata | suricatta | Herpestidae | 35470 | F | van Staaden 1994; Iwaniuk et al. 1999 | ||
39 | Sc | Suricata | suricatta | Herpestidae | 35469 | M | van Staaden 1994; Iwaniuk et al. 1999 | ||
40 | Sf | Mephitis | mephitis | Mephitidae | 35951 | F | Wade-Smith and Verts 1982; Van Valkenburgh 1987; Samuels and Van Valkenburgh 2008 | ||
41 | Sf | Mephitis | mephitis | Mephitidae | 39321 | M | Wade-Smith and Verts 1982; Van Valkenburgh 1987; Samuels and Van Valkenburgh 2008 | ||
42 | Sa | Lontra | canadensis | Mustelidae | 32226 | F | Gingerich 2003 | ||
43 | Sa | Lontra | canadensis | Mustelidae | 32230 | M | Gingerich 2003 | ||
44 | Sa | Mustela | vison | Mustelidae | 41780 | F | Howell 1930; Larivié́re 1999; Gingerich 2003 | ||
45 | Sa | Mustela | vison | Mustelidae | 35223 | M | Howell 1930; Larivié́re 1999; Gingerich 2003 | ||
46 | Sc | Gulo | gulo | Mustelidae | 14200 | F | Van Valkenburgh 1987; Pasitschniak-Arts and Larivié́re 1995 | ||
47 | Sc | Gulo | gulo | Mustelidae | 34417 | M | Van Valkenburgh 1987; Pasitschniak-Arts and Lariviére 1995 | ||
48 | Sf | Spilogale | putorius | Mustelidae | 39331 | F | Heinrich and Houde 2006 | ||
49 | Sf | Spilogale | putorius | Mustelidae | 39155 | M | Heinrich and Houde 2006 | ||
50 | T | Mustela | erminea | Mustelidae | 39366 | F | Nowak 1999 | ||
51 | T | Mustela | erminea | Mustelidae | 72863 | M | Nowak 1999 | ||
52 | T | Mustela | putorius | Mustelidae | 32599 | F | Nowak 1999 | ||
53 | T | Mustela | putorius | Mustelidae | 58727 | M | Nowak 1999 | ||
54 | A | Paguma | larvata | Viverridae | 73281 | U | Nowak 1999 | ||
Cingulata | |||||||||
55 | F | Cabassous | centralis | Dasypodidae | 34167 | F | Nowak 1999 | ||
56 | F | Dasypus | novemcinctus | Dasypodidae | 20735 | F | Samuels and Van Valkenburgh 2008 | ||
57 | F | Dasypus | novemcinctus | Dasypodidae | 22458 | M | Samuels and Van Valkenburgh 2008 | ||
58 | F | Euphractus | sexcinctus | Dasypodidae | 35468 | M | Redford and Wetzel 1985; Nowak 1999 | ||
Eulipotyphla | |||||||||
59 | T | Echinosorex | gymnurus | Erinaceidae | S2008 | U | Nowak 1999; Salton and Sargis 2008 | ||
60 | Sf | Solenodon | paradoxus | Solenodontidae | S2008 | U | Nowak 1999; Salton and Sargis 2008 | ||
Lagomorpha | |||||||||
61 | S | Brachylagus | idahoensis | Leporidae | 38628 | F | Nowak 1999 | ||
62 | S | Brachylagus | idahoensis | Leporidae | 38631 | M | Nowak 1999 | ||
63 | S | Lepus | americanus | Leporidae | 21111 | F | Nowak 1999 | ||
64 | S | Lepus | americanus | Leporidae | 33263 | M | Nowak 1999 | ||
65 | Sf | Ochotona | princeps | Ochtonidae | 18435 | F | Nowak 1999 | ||
66 | Sf | Ochotona | princeps | Ochtonidae | 60068 | M | Nowak 1999 | ||
Macroscelidea | |||||||||
67 | S | Petrodromus | tetradactylus | Macroscelididae | S2008 | U | Salton and Sargis 2008 | ||
68 | T | Elephantulus | rufescens | Macroscelididae | 35475 | F | Nowak 1999 | ||
69 | T | Elephantulus | rufescens | Macroscelididae | 34189 | M | Nowak 1999 | ||
Primates | |||||||||
70 | A | Callithrix | pygmaea | Cebidae | 39003 | F | Nowak 1999 | ||
71 | A | Callithrix | pygmaea | Cebidae | 39005 | M | Nowak 1999 | ||
72 | A | Leontopithecus | rosalia | Cebidae | 75541 | F | Nowak 1999 | ||
73 | A | Saguinus | oedipus | Cebidae | 35406 | F | Nyakatura et al. 2008 | ||
74 | A | Saguinus | oedipus | Cebidae | 35405 | M | Nyakatura et al. 2008 | ||
75 | A | Saimiri | sciureus | Cebidae | 82302 | F | Nowak 1999 | ||
76 | A | Saimiri | sciureus | Cebidae | 39014 | M | Nowak 1999 | ||
Rodentia | |||||||||
77 | F | Aplodontia | rufa | Aplodontiidae | 34071 | F | Samuels and Van Valkenburgh 2008 | ||
78 | F | Aplodontia | rufa | Aplodontiidae | 34058 | M | Samuels and Van Valkenburgh 2008 | ||
79 | Sa | Castor | canadensis | Castoridae | 34116 | F | Nowak 1999 | ||
80 | Sa | Castor | canadensis | Castoridae | 34588 | M | Nowak 1999 | ||
81 | T | Cavia | porcellus | Caviidae | 72830 | F | Nowak 1999 | ||
82 | T | Cavia | porcellus | Caviidae | 72831 | M | Nowak 1999 | ||
83 | F | Microtus | pennsylvanicus | Cricetidae | 76622 | F | Nowak 1999 | ||
84 | F | Microtus | pennsylvanicus | Cricetidae | 34326 | M | Nowak 1999 | ||
85 | Sa | Ondatra | zibethicus | Cricetidae | 34324 | F | Howell 1930; Gingerich 2003 | ||
86 | Sa | Ondatra | zibethicus | Cricetidae | 72879 | M | Howell 1930; Gingerich 2003 | ||
87 | Sf | Synaptomys | borealis | Cricetidae | 66559 | F | Nowak 1999 | ||
88 | Sf | Synaptomys | borealis | Cricetidae | 66561 | M | Nowak 1999 | ||
89 | S | Zapus | princeps | Dipodidae | 76721 | F | Nowak 1999 | ||
90 | S | Zapus | princeps | Dipodidae | 75136 | M | Nowak 1999 | ||
91 | Sc | Coendou | prehensilis | Erethizontidae | 34176 | F | Nowak 1999 | ||
92 | F | Thomomys | bottae | Geomyidae | 44624 | F | Nowak 1999 | ||
93 | F | Thomomys | bottae | Geomyidae | 38261 | M | Nowak 1999 | ||
94 | F | Chaetodipus | fallax | Heteromyidae | 47378 | F | Lackey 1996 | ||
95 | F | Chaetodipus | fallax | Heteromyidae | 47379 | M | Lackey 1996 | ||
96 | S | Dipodomys | deserti | Heteromyidae | 78740 | F | Samuels and Van Valkenburgh 2008 | ||
97 | Sf | Octodon | degus | Octodontidae | 48984 | F | Nowak 1999 | ||
98 | Sf | Octodon | degus | Octodontidae | 39501 | M | Nowak 1999 | ||
99 | A | Sciurus | aberti | Sciuridae | 35278 | F | Nowak 1999 | ||
100 | A | Sciurus | aberti | Sciuridae | 82349 | M | Nowak 1999 | ||
101 | A | Sciurus | carolinensis | Sciuridae | 20646 | F | Nowak 1999 | ||
102 | A | Sciurus | carolinensis | Sciuridae | 30010 | M | Nowak 1999 | ||
103 | G | Glaucomys | sabrinus | Sciuridae | 36334 | F | Samuels and Van Valkenburgh 2008 | ||
104 | G | Glaucomys | sabrinus | Sciuridae | 35129 | M | Samuels and Van Valkenburgh 2008 | ||
105 | G | Glaucomys | volans | Sciuridae | 43897 | F | Nowak 1999 | ||
106 | G | Glaucomys | volans | Sciuridae | 32254 | M | Nowak 1999 | ||
107 | Sf | Cynomys | ludovicianus | Sciuridae | 75774 | M | Nowak 1999 | ||
Scandentia | |||||||||
108 | Sc | Tupaia | glis | Tupaiidae | 34227 | F | Van Valkenburgh 1987 | ||
109 | Sc | Tupaia | glis | Tupaiidae | 34225 | M | Van Valkenburgh 1987 |
TABLE 5. Landmark number, location, type, and source reference for two-dimensional geometric morphometrics (2D GM) points on therian distal humeri. All points were digitized on distal images of specimens; see Figure 3 for more details. Numbers in parentheses are the landmark number for the point within the referenced source. Abbreviations: LM, landmark; Type, landmark type according to Zelditch et al. 2004.
LM | Location | Type | Source Reference |
1 | Lateral point on the posterior trochlea | II | Steiner-Souza et al. 2010 |
2 | Medial point on the posterior trochlea | II | Steiner-Souza et al. 2010 |
3 | Medial point on the anterior trochlea | II | Steiner-Souza et al. 2010 |
4 | Lateral point on the anterior capitulum | II | Steiner-Souza et al. 2010 |
5 | Most lateral point on the ectepicondyle | III | Schutz and Guralnick 2007 (5); Steiner-Souza et al. 2010 |
6 | Most medial point on the entepicondyle | III | Milne et al. 2009 (12); Steiner-Souza et al. 2010 |