ACKNOWLEDGMENTS

We thank S. Wallace, J. Mead, and T. Laughlin for comments and suggestions on an earlier version of this manuscript.

Thanks to the prep lab (B. Compton and S. Haugrud) and collections (B. Woodward and A. Nye) staff at the ETSU and General Shale Brick Natural History Museum at the Gray Fossil Site for preparing and providing access to the fossil specimens examined. W. Devender, D. Parmley, the New Jersey State Museum, and Carnegie Museum of Natural History loaned comparative specimens for this study. C. Bell and J. Olori at UT Austin provided reprints and helpful comments during the initial phases of this study.

Financial support was provided by the ETSU Research and Development Committee (to B.W. Schubert), and a Graduate Council Research Grant, and an NSF GK-12 Fellowship (National Science Foundation, Division of Graduate Education GK-12 Program, grant number DGE-074236) to G.S. Boardman.